Chamaecyparis lawsoniana
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Chamaecyparis lawsoniana, known as Port Orford cedar<ref name=PLANTS>Template:PLANTS</ref> or Lawson's cypress,<ref name=BSBI07>Template:BSBI 2007</ref> is a species of conifer in the genus Chamaecyparis, family Cupressaceae. It is native to Oregon and northwestern California, and grows from sea level up to Template:Convert in the valleys of the Klamath Mountains, often along streams.
Description
A large evergreen tree, specimens mature up to Template:Convert tall or more (exceptionally to Template:Convert),<ref name="GD">Template:Cite web</ref> with trunks Template:Convert in diameter, exceptionally Template:Convert.<ref name="GD"/><ref name=":02">Template:Cite book</ref> The bark is silver-brown, vertically furrowed, and Template:Convert thick near the base.<ref name=":02" /> The foliage is arranged in lacy, flat sprays with a feathery appearance,<ref name=":02" /> usually somewhat glaucous (i.e. blue-green) in color. The leaves are scale-like, Template:Convert long, with narrow white markings on the underside, and produced on somewhat flattened shoots. The foliage gives off a rather pungent scent, not unlike parsley. The seed cones are globose, Template:Convert in diameter, with 6–10 scales, green at first, maturing brown in early fall, 6–8 months after pollination. The male cones are Template:Convert long, dark red, turning brown after pollen release in early spring. The seeds fall quickly and can float on water.<ref name=":02" />
Old specimens lack branches near the base and often have dead tops. They can live up to about 600 years of age.<ref name=":02" />
Taxonomy
The species was first documented near Port Orford, Oregon, and introduced into cultivation in 1854 by collectors working for Charles Lawson FRSE<ref>Template:Cite book</ref> of the Lawson & Son nursery in Edinburgh, Scotland, after whom it was named as Lawson's Cypress by the describing botanist Andrew Murray. The United States Department of Agriculture officially calls it by the name Port Orford cedar,<ref name="PLANTS" /> as do most people in its native area, but some botanists prefer to use the name Lawson's cypress (or in very rare instances Port Orford cypress) instead. The name "Lawson's cypress" is widely used in horticulture.
Distribution and habitat
The species is native to southwestern Oregon and northwestern California, and grows from sea level up to Template:Convert<ref name=":02" /> in the valleys of the Klamath Mountains, often along streams. It fares best at the north end of its range.<ref name=":02" />
Ecology
The thick bark provides resistance to wildfires, and the species regenerates well on disrupted land in a variety of soils, but requires consistent moisture. It is shade tolerant, but not as much so as competing species western hemlock and white fir.<ref name=":02" /> The old-growth population near Coos Bay, Oregon, was wiped out by logging and wildfires in 1867–1868, and again by fire and root disease in 1936.<ref name=":02" />
Disease
In the wild, the species is seriously and uniquely threatened by a root disease caused by the oomycete pathogen, Phytophthora lateralis, accidentally introduced in the early 1950s following the fungus's arrival to the Pacific Northwest on nursery trees.<ref name=":02" /> This disease is also a problem for horticultural plantings in some parts of North America. The tree is sometimes killed, though less often, by other species of Phytophthora.Template:Cn
Phytophthora lateralis infection begins when mycelium, from a germinated spore, invade the roots. The infection then spreads through the inner bark and cambium around the base of the tree. Spread up the trunk is generally limited. Infected tissue dies and effectively girdles the tree. Large trees are more likely to be infected than small trees due to larger root areas (although all trees at the edges of infected streams will eventually succumb). However, large trees can often live with the infections for a longer duration (up to several years).Template:Cn
C. lawsoniana in streamside populations are highly susceptible to P. lateralis infection. However, the rate of fungal spread through populations in dry upland areas appears to be slow. P. lateralis spreads through water via mobile spores (zoospores). The fungus also produces resting spores (chlamydospores) that can persist in soil for a long period of time. New infections generally begin when soil is transferred from an infected population to a non-infected population via water, human or animal movement.<ref name=":02" /> After initial infection in streamside populations, secondary spread via zoospores quickly infects all downstream individuals.Template:Cn
Human facilitated spread is thought to be responsible for most new, and all long-distance, infections. Soil on vehicle tires, especially logging trucks and other off-road vehicles, is considered the most pressing problem due to the volume of soil that can be carried and the traffic rate in and between susceptible areas. Spread on boots and mountain bike tires has also been suggested and probably contributes to new infections locally.<ref>Template:Cite news</ref> Animal-facilitated spread is thought to occur, but is localized.
The United States Forest Service and Bureau of Land Management attempt to prevent Phytophthora spread through road closures, monitoring, research and education. Research has focused on determining the dynamics and mechanisms of spread, as well as attempts to breed resistant trees. One solution against Phytophtera is known generically as Mancozeb and also commercially known as Dithane (C). Commercial preparations of the parasitic fungus Pythium oligandrum are licensed for pest control, and documented to predate many species of Phytophthora.<ref>Template:Cite web</ref>
Similar species
The extinct Eocene species Chamaecyparis eureka, known from fossils found on Axel Heiberg Island in Canada, is noted as resembling C. lawsoniana and C. pisifera.<ref name="Kotyk2003">Template:Cite journal</ref>
The associated genus Calocedrus (incense-cedar) has thick orange-brown bark and the bark of Thuja plicata (western redcedar) is comparatively thin; both have different foliage than Port Orford cedar.<ref name=":02" />
Cultivation
Chamaecyparis lawsoniana thrives best in well-drained but moist soils, in a fairly sheltered position in full sun. Several hundred named cultivars of varying crown shape, growth rates and foliage color have been selected for planting in parks and gardens. In the United Kingdom (UK) the following have gained the Royal Horticultural Society's Award of Garden Merit (confirmed 2017):<ref>Template:Cite web</ref>
| Name | Height (m) | Spread (m) | Shape | Foliage Colour | Ref. |
|---|---|---|---|---|---|
| 'Aurea Densa' | 1.5 | 1.0 | broadly conical | yellow-green | <ref>Template:Cite web</ref> |
| 'Chilworth Silver' | 2.5 | 1.0 | broadly columnar | silver-blue | <ref>Template:Cite web</ref> |
| 'Dik's Weeping' | 12.0 | 4.0 | columnar, weeping | blue-green | <ref>Template:Cite web</ref> |
| 'Ellwoodii' | 8.0 | 4.0 | columnar | grey-green | <ref>Template:Cite web</ref> |
| 'Ellwood's Gold' | 4.0 | 1.0 | columnar | yellow/grey-green | <ref>Template:Cite web</ref> |
| 'Ellwood's Pillar' | 2.5 | 1.5 | narrowly columnar | grey-green | <ref>Template:Cite web</ref> |
| 'Fletcheri' | 12.0 | 4.0 | columnar | grey-green | <ref>Template:Cite web</ref> |
| 'Gimbornii' | 2.5 | 2.5 | rounded | grey-green | <ref>Template:Cite web</ref> |
| 'Golden Wonder' | 20.0 | 4.0 | columnar | yellow-green | <ref>Template:Cite web</ref> |
| 'Grayswood Feather' | 4.0 | 1.5 | narrowly columnar | green | <ref>Template:Cite web</ref> |
| 'Green Globe' | 0.3 | 0.3 | rounded | dark green | <ref>Template:Cite web</ref> |
| 'Imbricata Pendula' | 8.0 | 4.0 | conical, weeping, pendulous | green | <ref>Template:Cite web</ref> |
| 'Kilmacurragh' | 12.0 | 4.0 | columnar | dark green | <ref>Template:Cite web</ref> |
| 'Lanei Aurea' | 8.0 | 2.5 | columnar | yellow-green | <ref>Template:Cite web</ref> |
| 'Little Spire' | 2.5 | 1.5 | conical | dark green | <ref>Template:Cite web</ref> |
| 'Minima Aurea' | 1.0 | 1.0 | conical | yellow-green | <ref>Template:Cite web</ref> |
| 'Minima Glauca' | 2.5 | 2.5 | rounded | blue-green | <ref>Template:Cite web</ref> |
| 'Pembury Blue' | 12.00+ | 8.00 | conical | blue-green | <ref>Template:Cite web</ref> |
| 'Pygmaea Argentea' | 2.5 | 1.0 | rounded | cream/blue-green | <ref>Template:Cite web</ref> |
| 'Stardust' | 4.0 | 1.5 | columnar | yellow-green | <ref>Template:Cite web</ref> |
| 'Summer Snow' | 2.5 | 1.5 | conical | cream/green | <ref>Template:Cite web</ref> |
| 'Triomf van Boskoop' | 25.0 | 8.0 | broadly columnar | grey-green | <ref>Template:Cite web</ref> |
| 'Wisselii' | 12.0+ | 4.0 | narrowly conical | blue-green | <ref>Template:Cite web</ref> |
| 'Wissel's Saguaro' | 6.0 | 1.0 | narrow, upright | blue-green | <ref>Template:Cite web</ref> |
Uses
The species was discovered by Euro-Americans in the 1850s. The wood is light yet has great strength and rot resistance, even after long exposure to salt water.<ref name=":02" /> Its properties resemble those of yellow-cedar, but was historically more available in the region.<ref name=":02" /> On shores lacking docks, logs were transported via high-line cable directly onto ship decks. It was valued for boatbuilding.<ref name=":02" /> The species was important to Oregon's lumber industry until the 1950s when it was crippled by disease.<ref name=":02" /> It was preferred for storage battery cell separation, Venetian blinds, and other uses.<ref name=":02" /> Quality specimens eventually began to be shipped almost exclusively to East Asia, where it is highly valued.<ref name=":02" /> Large amounts have been exported to Japan where it is used in making coffins, shrines, and temples.<ref>Snyder 1999 pg. 225</ref> Its lumber is known for its highly fragrant ginger aroma, caused by an oil which repulses decay and insects, including termites; this oil has been used as an insecticide.<ref name=":02" /> Due to the straightness of its grain, it is also one of the preferred woods for the manufacture of arrow shafts. It is also considered an acceptable, though not ideal, wood for construction of aircraft.<ref>Kroes, Watkins, and Delp: Aircraft Maintenance and Repair, sixth edition, page 66, McGraw Hill, 1993</ref>
The wood is considered more than acceptable for use in stringed instruments. Its fine grain, good strength and tonal quality are highly regarded for soundboards in guitar making.<ref>Template:Cite web</ref>
Gallery
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Foliage with cones: immature seed cones below, pollen cones above
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Male cones in spring with diagnostic red colour
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Mature female cones
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Cultivar 'Imbricata Pendula', Rosemoor, Devon, UK
See also
References
- Snyder, Gary. 1999. "The Gary Snyder Reader". Counterpoint. Template:ISBN
Further reading
- Template:Cite iucn Listed as Vulnerable (VU A1de+2e v2.3)
- Siskiyou National Forest has posted precautions for persons entering areas with Port Orford Cedar populations ([1]).
- Template:Cite journal
- Hunt, J. 1959. Phytophthora lateralis on Port-Orford-cedar. Research Note 172: 1–6. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station.
- Template:Cite journal
- Torgeson, D. C., Young, R. A., & Milbrath, J. A. 1954. Phytophthora root rot diseases of Lawson cypress and other ornamentals. Oregon Agricultural Experiment Station Bulletin. 537: 1–18. Corvallis, OR: Oregon State College.
- Template:Cite journal
- Template:Cite journal
- Zobel, D. B., Roth, L. F., & Hawk, G. M. 1985. Ecology, pathology, and management of Chamaecyparis lawsoniana. General Technical Report. PNW-184: 1–161. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Forest and Range Experiment Station.
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External links
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