View source for Thomas Henry Huxley

{{Short description|English biologist and comparative anatomist (1825–1895)}}
{{Redirect|Thomas Huxley|the Lieutenant-Colonel|Thomas Huxley (British Army officer)}}
{{Use British English|date=February 2024}}
{{Use dmy dates|date=February 2024}}
{{Infobox scientist
| honorific_prefix  = [[The Right Honourable]]
| name              = Thomas Henry Huxley
| honorific_suffix  = {{post-nominals|country=GBR|size=100%|FRS|FLS|HFRSE}}
| image             = Thomas Henry Huxley. Photograph by Lock & Whitfield. Wellcome V0026596.jpg
| caption           = [[Woodburytype]] print of Huxley (1880 or earlier)
| birth_date        = {{birth date|1825|5|4|df=y}}
| birth_place       = [[Ealing]], London, England
| death_date        = {{death date and age|1895|6|29|1825|5|4|df=y}}
| death_place       = [[Eastbourne]], Sussex, England
| education         = {{plainlist|
* Sydenham College, London<ref>Adrian J. Desmond, ''Huxley: From Devil's Disciple to Evolution's High Priest'', Addison-Wesley, 1994, 1915, p. 651 n. 8.</ref>
* [[Charing Cross Hospital]]}}
| academic_advisors = [[Thomas Wharton Jones]]
| notable_students  = {{plainlist|
* [[Michael Foster (physiologist)|Michael Foster]]
* [[H. G. Wells]]}}
| known_for         = [[Evolution]], [[science education]], [[agnosticism]]
| awards            = {{plainlist|
* [[Royal Medal]] {{small|(1852)}}
* [[Wollaston Medal]] {{small|(1876)}}
* [[Clarke Medal]] {{small|(1880)}}
* [[Copley Medal]] {{small|(1888)}}
* [[Linnean Medal]] {{small|(1890)}}
* [[Hayden Memorial Geological Award]] {{small|(1893)}}}}
| children = Eight, including [[Marian Huxley]] and [[Leonard Huxley (writer)|Leonard Huxley]]
| signature         = Thomas Huxley Signature in Cassell's Universal Portrait Gallery.svg
| field             = [[Zoology]]; [[comparative anatomy]]
| work_institutions = [[Royal Navy]], [[Royal College of Surgeons of England|Royal College of Surgeons]], [[Royal School of Mines]], [[Royal Institution]] [[University of London]]
}}

'''Thomas Henry Huxley''' (4 May 1825 – 29 June 1895) was an English [[biologist]] and [[anthropologist]] who specialised in [[comparative anatomy]]. He has become known as "'''Darwin's Bulldog'''" for his advocacy of [[Charles Darwin]]'s theory of [[evolution]].<ref>{{Harvnb|Encyclopædia Britannica Online|2006}}</ref>

The stories regarding Huxley's famous [[1860 Oxford evolution debate]] with [[Samuel Wilberforce]] were a key moment in the wider acceptance of [[evolution]] and in his own career, although some historians think that aspects of the surviving story of the debate are a later fabrication.<ref>Livingstone, David. "Myth 17. That Huxley Defeated Wilberforce in Their Debate over Evolution and Religion," in Numbers, Ronald L., ed. Galileo goes to jail and other myths about science and religion. No. 74. Harvard University Press, 2009, 152–160.</ref> Huxley had been planning to leave [[Oxford]] on the previous day, but, after an encounter with [[Robert Chambers (journalist)|Robert Chambers]], the author of ''[[Vestiges of the Natural History of Creation|Vestiges]]'', he changed his mind and decided to join the debate. Wilberforce was coached by [[Richard Owen]], against whom Huxley also debated about whether humans were closely related to apes.

Huxley was slow to accept some of Darwin's ideas, such as [[gradualism]], and was undecided about [[natural selection]], but despite this, he was wholehearted in his public support of Darwin. Instrumental in developing scientific [[education]] in Britain, he fought against the more extreme versions of religious tradition. Huxley coined the term "[[agnosticism]]" in 1869 and elaborated on it in 1889 to frame the nature of claims in terms of what is knowable and what is not.

Huxley had little formal schooling and was virtually self-taught. According to Edward Poulton, he became perhaps the finest [[comparative anatomy|comparative anatomist]] of the later 19th century.<ref>Poulton E. B. 1909. Charles Darwin and the origin of species. London.</ref> He worked on [[invertebrates]], clarifying relationships between groups previously little understood. Later, he worked on [[vertebrates]], especially on the relationship between apes and humans. After comparing ''[[Archaeopteryx]]'' with ''[[Compsognathus]]'', he concluded that [[Evolution of birds|birds]] evolved from small carnivorous [[Theropoda|dinosaurs]], a view now held by modern biologists.

The tendency has been for this fine anatomical work to be overshadowed by his energetic and controversial activity in favour of evolution, and by his extensive public work on scientific education, both of which had significant effects on society in Britain and elsewhere.<ref>[[Ray Lankester|Lankester, Ray]] 1895. The Right Hon. T. H. Huxley. ''Athenaeum'', 6 July. Lankester commented that Huxley was "only accidentally a zoologist".</ref><ref>{{Harvnb|Desmond|1997}}. 'Huxley in perspective', 235–261, an outstanding summary of Huxley in his social & historical context, scarcely mentions his zoological work.</ref> Huxley's 1893 [[Romanes Lecture]], "''[[s:Evolution and Ethics|Evolution and Ethics]]''", is exceedingly influential in China; the Chinese translation of Huxley's lecture even transformed the Chinese translation of Darwin's ''Origin of Species''.<ref>{{cite journal |last=Jin |first=Xiaoxing |title=Translation and transmutation: the Origin of Species in China |journal=The British Journal for the History of Science |volume=52 |issue=1 |pages=117–141 |year=2019 |pmid=30587253 |s2cid=58605626 |doi=10.1017/S0007087418000808}}</ref>

== Early life ==
{{tone|section|date=June 2022}}
{{more citations needed|section|date=June 2022}}
Thomas Henry Huxley was born in [[Ealing]], then a village in [[Middlesex]]. He was the second youngest of eight children<ref>{{cite web |title=Thomas Huxley |url=https://ucmp.berkeley.edu/history/thuxley.html |access-date=2023-02-26 |website=ucmp.berkeley.edu |archive-date=26 February 2023 |archive-url=https://web.archive.org/web/20230226054420/https://ucmp.berkeley.edu/history/thuxley.html |url-status=live}}</ref> of George Huxley and Rachel Withers. His parents were members of the Church of England, but he sympathized with [[Nonconformist (Protestantism)|nonconformists]].<ref name="Brit"/> Like some other British scientists of the nineteenth century such as [[Alfred Russel Wallace]], Huxley was brought up in a literate middle-class family which had fallen on hard times. His father was a mathematics [[schoolmaster]] at [[Great Ealing School]] until it closed,<ref>Bibby, amongst others, queried this account, which owes its origin to Leonard Huxley's biography (1900). Bibby, Cyril. 1959. ''T. H. Huxley: scientist, humanist and educator''. Watts, London. p. 3–4</ref> putting the family into financial difficulties. As a result, Thomas left school at the age of 10, after only two years of formal schooling.<ref name="Brit"/>

Despite this lack of formal schooling, Huxley was determined to educate himself. He became one of the great [[autodidact]]s of the nineteenth century. At first he read [[Thomas Carlyle]], [[James Hutton]]'s ''Geology'', and [[Sir William Hamilton, 9th Baronet|Sir William Hamilton]]'s ''Logic''. In his teens, he taught himself [[German language|German]], eventually becoming fluent and used by [[Charles Darwin]] as a translator of scientific material in German. He learned [[Latin language|Latin]], and enough [[Greek language|Greek]] to read Aristotle in the original.<ref>{{cite news |title=The Project Gutenberg eBook of Thomas Henry Huxley - A Sketch of His Life and Work by P. Chalmers Mitchell, M.A. (Oxon.). |url=https://www.gutenberg.org/files/16935/16935-h/16935-h.htm#p186 |access-date=11 January 2023 |archive-date=11 January 2023 |archive-url=https://web.archive.org/web/20230111132516/https://www.gutenberg.org/files/16935/16935-h/16935-h.htm#p186 |url-status=live}}</ref>

[[Image:Young Huxley RN.jpg|thumb|Huxley, aged 21]]

Later on, as a young adult, he made himself an expert, first teaching himself about [[invertebrates]], and later on [[vertebrates]]. He did many of the illustrations for his publications on marine invertebrates. In his later debates and writing on science and religion, his grasp of theology was better than many of his clerical opponents.<ref name=autogenerated3>{{Harvnb|Desmond|1994}}</ref><ref name=autogenerated4>{{Harvnb|Huxley|1900}}</ref>

He was apprenticed for short periods to several medical practitioners: at 13 to his brother-in-law John Cooke in Coventry, who passed him on to Thomas Chandler, notable for his experiments using [[animal magnetism|mesmerism]] for medical purposes. Chandler's practice was in London's [[Rotherhithe]] amidst the squalor endured by the [[Charles Dickens|Dickens]]ian poor.<ref>Chesney, Kellow 1970. ''The Victorian underworld''. Temple Smith, London; Pelican 1972, pp. 105, 421.</ref> Afterward, another brother-in-law took him on: John Salt, his eldest sister's husband. Now aged 16, Huxley entered Sydenham College (behind [[University College Hospital]]), a cut-price anatomy school. All this time Huxley continued his programme of reading, which more than made up for his lack of formal schooling.

A year later, buoyed by excellent results and a silver medal prize in the [[Society of Apothecaries|Apothecaries']] yearly competition, Huxley was admitted to study at [[Charing Cross Hospital]], where he obtained a small scholarship. At Charing Cross, he was taught by [[Thomas Wharton Jones]], Professor of Ophthalmic Medicine and Surgery at [[University College London]]. Jones had been [[Robert Knox (surgeon)|Robert Knox]]'s assistant when Knox bought cadavers from [[Burke and Hare]].<ref>The cut-price anatomy schools and Robert Knox are well treated in Desmond's account of materialist medical dissidents of the 1820s and 30s: Desmond A. 1989. ''The politics of evolution: morphology, medicine and reform in radical London''. Chicago.</ref> The young Wharton Jones, who acted as go-between, was exonerated of crime, but thought it best to leave Scotland. In 1845, under Wharton Jones' guidance, Huxley published his first scientific paper demonstrating the existence of a hitherto unrecognised layer in the inner sheath of hairs, a layer that has been known since as [[Huxley's layer]]. Later in life, Huxley organised a pension for his old mentor.

At twenty he passed his First M.B. examination at the [[University of London]], winning the gold medal for [[anatomy]] and [[physiology]]. However, he did not present himself for the final (Second M.B.) exams and consequently did not qualify for a university degree. His apprenticeships and exam results formed a sufficient basis for his application to the [[Royal Navy]].<ref name=autogenerated3 /><ref name=autogenerated4 />

== Voyage of the ''Rattlesnake'' ==
Aged 20, Huxley was too young to apply to the [[Royal College of Surgeons of England|Royal College of Surgeons]] for a licence to practise, yet he was "deep in debt".<ref>{{Harvnb|Desmond|1994|p=35}}</ref> So, at a friend's suggestion, he applied for an appointment in the Royal Navy. He had references on character and certificates showing the time spent on his apprenticeship and on requirements such as dissection and pharmacy. [[William Burnett (physician)|William Burnett]], the Physician General of the Navy, interviewed him and arranged for the College of Surgeons to test his competence (by means of a ''[[Oral exam|viva voce]]'')<!-- viva voce links to a disambiguation page, but the best Wikipedia choice might be [[Thesis#Thesis examinations]] which would confuse the reader even more ("often an oral examination" - it's always oral); the wiktionary link alternative is worse in it makes the reader think of a dental procedure. Leave this ambiguous for now, until a UK-specific explanation arises. -->.{{Citation needed|date=April 2023}}

[[File:Rattlesnake2.jpg|thumb|left|upright|[[HMS Rattlesnake (1822)|HMS ''Rattlesnake'']] by the ship's artist [[Oswald Brierly]]]]

Finally, Huxley was made [[surgery|Assistant Surgeon]] ('surgeon's mate', but in practice marine naturalist) to [[HMS Rattlesnake (1822)|HMS ''Rattlesnake'']], about to set sail on a voyage of discovery and surveying to New Guinea and Australia. The ''Rattlesnake'' left England on 3 December 1846 and, once it arrived in the southern hemisphere, Huxley devoted his time to the study of marine invertebrates.<ref>{{Harvnb|Huxley|1935}}</ref> He began to send details of his discoveries back to England, where publication was arranged by [[Edward Forbes]] FRS (who had also been a pupil of Knox). Both before and after the voyage Forbes was something of a mentor to Huxley.{{Citation needed|date=April 2023}}

Huxley's paper "On the anatomy and the affinities of the family of Medusae" was published in 1849 by the [[Royal Society]] in its ''Philosophical Transactions''. Huxley united the Hydroid and Sertularian polyps with the Medusae to form a class to which he subsequently gave the name of ''[[Hydrozoa]]''. The connection he made was that all the members of the class consisted of two cell layers, enclosing a central cavity or stomach. This is characteristic of the [[phylum]] now called the ''[[Cnidaria]]''. He compared this feature to the serous and mucous structures of embryos of higher animals. When at last he got a grant from the Royal Society for the printing of plates, Huxley was able to summarise this work in ''The Oceanic Hydrozoa'', published by the [[Ray Society]] in 1859.<ref>{{Harvnb|Di Gregorio|1984}}</ref><ref>{{Harvnb|Huxley|1984}}</ref>

[[File:Australian woman by Thomas Henry Huxley.jpg|thumb|upright=0.6|Australian woman: Pencil drawing by Huxley]]

The value of Huxley's work was recognised and, on returning to England in 1850, he was elected a [[Fellow of the Royal Society]]. In the following year, at the age of twenty-six, he not only received the Royal Society Medal but was also elected to the Council. He met [[Joseph Dalton Hooker]] and [[John Tyndall]],<ref>{{Harvnb|Tyndall|1896|Ref=CITEREFTyndall1896|}} pp. 7, 9, 66, 71.</ref> who remained his lifelong friends. The Admiralty retained him as a nominal assistant-surgeon, so he might work on the specimens he collected and the observations he made during the voyage of the ''Rattlesnake''. He solved the problem of ''[[Larvacea|Appendicularia]]'', whose place in the animal kingdom [[Johannes Peter Müller]] had found himself wholly unable to assign. It and the [[Ascidiacea|Ascidians]] are both, as Huxley showed, [[tunicates]], today regarded as a sister group to the vertebrates in the phylum ''[[Chordata]]''.<ref>{{Harvnb|Holland|2007|}} pp. 153–5</ref> Other papers on the [[morphology (biology)|morphology]] of the [[cephalopod]]s and on [[brachiopod]]s and [[rotifer]]s are also noteworthy.<ref name=autogenerated3 /><ref name=autogenerated4 /><ref name="foster1903">{{Harvnb|Foster|Lankester|2007}}</ref>

== Later life ==
Following a lecture at the Royal Institution on 30 April 1852 Huxley indicated that it remained difficult to earn a living as a scientist alone. This was demonstrated in a letter written on 3 May 1852, where he states "Science in England does everything—but PAY. You may earn praise but not pudding".<ref>{{cite web |date=December 7, 2023 |title=Life and Letters of Thomas Henry Huxley — Volume 1 |url=https://www.gutenberg.org/cache/epub/5084/pg5084.html |access-date=December 7, 2023 |website=Project Gutenberg}}</ref> However, Huxley effectively resigned from the navy by refusing to return to active service, and in July 1854 he became professor of natural history at the [[Royal School of Mines]] and naturalist to the [[British Geological Survey]] in the following year. In addition, he was Fullerian Professor at the [[Royal Institution]] 1855–1858 and 1865–1867; Hunterian Professor at the [[Royal College of Surgeons of England|Royal College of Surgeons]] 1863–1869; president of the [[British Association for the Advancement of Science]] 1869–1870; president of the [[Quekett Microscopical Club]] 1878; president of the [[Royal Society]] 1883–1885; Inspector of Fisheries 1881–1885; and president of the [[Marine Biological Association of the United Kingdom|Marine Biological Association]] 1884–1890.<ref name=autogenerated4/> He was elected as a member to the [[American Philosophical Society]] in 1869.<ref>{{cite web |title=APS Member History |publisher=[[American Philosophical Society]] |website=amphilsoc.org |url=https://search.amphilsoc.org/memhist/search?year=1869;year-max=1869;smode=advanced;startDoc=21 |access-date=2021-04-26 |archive-date=26 April 2021 |archive-url=https://web.archive.org/web/20210426181651/https://search.amphilsoc.org/memhist/search?year=1869;year-max=1869;smode=advanced;startDoc=21 |url-status=live}}</ref>

The thirty-one years during which Huxley occupied the chair of natural history at the Royal School of Mines included work on vertebrate palaeontology and on many projects to advance the place of science in British life. Huxley retired in 1885, after a bout of depressive illness which started in 1884. He resigned the presidency of the Royal Society in mid-term, the Inspectorship of Fisheries, and his chair (as soon as he decently could) and took six months' leave. His pension was £1200 a year.<ref name="Brit">{{Cite encyclopedia |title=Thomas Henry Huxley |series=British biologist |encyclopedia=Encyclopedia Britannica |url=https://www.britannica.com/biography/Thomas-Henry-Huxley |access-date=4 February 2018 |archive-date=22 January 2019 |archive-url=https://web.archive.org/web/20190122013913/https://www.britannica.com/biography/Thomas-Henry-Huxley |url-status=live}}</ref>

[[File:4MarlboroughPlace.png|thumb|4 Marlborough Place, London]]
Huxley's London home, in which he wrote many of his works, was at 4&nbsp;[[Marlborough Place]], [[St John's Wood]]. The house was extended in the early 1870s to include a large drawing and dining room at which he held informal Sunday gatherings.<ref name=IlstLondNews-1895-07-06>{{cite periodical |title=Professor Huxley's homes |date=6 July 1895 |periodical=The Illustrated London News |volume=107 |page=3}}</ref>

In 1890, he moved from London to [[Eastbourne]], where he had purchased land in the Staveley Road upon which a house was built, 'Hodeslea', under the supervision of his son-in-law F.&nbsp;Waller.<ref name=IlstLondNews-1895-07-06/> Here Huxley edited the nine volumes of his ''Collected Essays''. In 1894 he heard of [[Eugene Dubois]]' discovery in Java of the remains of ''Pithecanthropus erectus'' (now known as ''[[Homo erectus]]'').

[[File:HodesleaEastbourne.png|thumb|Hodeslea, Staveley Road, Eastbourne]]

He died of a heart attack (after contracting influenza and pneumonia) in 1895 in Eastbourne, and was buried in London at [[East Finchley Cemetery]]. No invitations were sent out but two hundred people turned up for the funeral; they included [[Joseph Dalton Hooker]], [[William Henry Flower]], [[Mulford B. Foster]], [[Edwin Lankester]], [[Joseph Lister, 1st Baron Lister|Joseph Lister]], and apparently [[Henry James]]. The family plot had been purchased upon the death of his eldest son Noel, who died of [[scarlet fever]] in 1860; Huxley's wife Henrietta Anne {{née|Heathorn}} and son Noel are also buried there.<ref>{{Harvp|Desmond|1997|p=230}}</ref>
[[Image:Thomas Huxley grave.jpg|thumb|Huxley's grave in [[East Finchley Cemetery]] in North London]]

===Family===
Huxley and his wife had five daughters and three sons:

* Noel Huxley (1856–1860), died aged&nbsp;3.<ref>{{Harvp|Desmond|1994|p=230|loc=Chapter: The Nature of the Beast}}</ref><ref>{{Harvp|Desmond|1994|pags=285–286|loc=Chapter: Buttered Angels & Bellowing Apes}}</ref>
* Jessie Oriana Huxley (1856–1927), married architect Fred Waller in 1877.
* [[Marian Huxley]] (1859–1887), married artist [[John Collier (painter)|John Collier]] in 1879.
* [[Leonard Huxley (writer)|Leonard Huxley]] (1860–1933), married [[Julia Huxley|Julia Arnold]].
* Rachel Huxley (1862–1934), married civil engineer Alfred Eckersley in 1884.
* Henrietta (Nettie) Huxley (1863–1940), married Harold Roller, travelled Europe as a singer.
* Henry Huxley (1865–1946), became a fashionable general practitioner in London.
* Ethel Huxley (1866–1941) married artist John Collier (widower of sister) in 1889.

=== Public duties and awards ===
From 1870 onwards, Huxley was to some extent drawn away from scientific research by the claims of public duty. He served on eight [[Royal Commission]]s, from 1862 to 1884. From 1871 to 1880 he was a Secretary of the [[Royal Society]] and from 1883 to 1885 he was president. He was president of the [[Geological Society]] from 1868 to 1870. In 1870, he was president of the [[British Association]] at Liverpool and, in the same year was elected a member of the newly constituted [[London School Board]]. He was president of the [[Quekett Microscopical Club]] from 1877 to 1879. He was the leading person amongst those who reformed the Royal Society, persuaded government about science, and established scientific education in British schools and universities.<ref name=autogenerated5>{{Harvp|Bibby|1959}}</ref>

He was awarded the highest honours then open to British men of science. The [[Royal Society]], who had elected him as Fellow when he was 25 (1851), awarded him the [[Royal Medal]] the next year (1852), a year before Charles Darwin got the same award. He was the youngest biologist to receive such recognition. 
He was elected to Honorary membership of the [[Manchester Literary and Philosophical Society]]<ref>Memoirs and proceedings of the Manchester Literary & Philosophical Society FOURTH SERIES Eighth VOLUME 1894</ref> in 1872.
Then later in life he received the [[Copley Medal]] in 1888 and the [[Darwin Medal]] in 1894; the [[Geological Society]] awarded him the [[Wollaston Medal]] in 1876; the [[Linnean Society]] awarded him the [[Linnean Medal]] in 1890. There were many other elections and appointments to eminent scientific bodies; these and his many academic awards are listed in the ''Life and Letters''. He turned down many other appointments, notably the Linacre chair in zoology at Oxford and the Mastership of [[University College, Oxford]].<ref>{{Harvp|Bibby|1972}}</ref>

[[Image:T.H.Huxley-1880s.jpg|thumb|{{center|Huxley <br/>by [[Alexander Bassano]] {{circa|1883}} }}]]

In 1873 the King of Sweden made Huxley, Hooker and Tyndall Knights of the [[Order of the Polar Star]]: they could wear the insignia but not use the title in Britain.<ref>{{Harvp|Desmond|1998|p=431}}</ref> Huxley collected many honorary memberships of foreign societies, academic awards and honorary doctorates from Britain and Germany. He also became a foreign member of the [[Royal Netherlands Academy of Arts and Sciences]] in 1892.<ref>{{cite web |title=T.H. Huxley (1825–1895) |publisher=[[Royal Netherlands Academy of Arts and Sciences]] |url=http://www.dwc.knaw.nl/biografie/pmknaw/?pagetype=authorDetail&aId=PE00001052 |access-date=19 July 2015 |archive-date=21 July 2015 |archive-url=https://web.archive.org/web/20150721212646/http://www.dwc.knaw.nl/biografie/pmknaw/?pagetype=authorDetail&aId=PE00001052 |url-status=live }}</ref> In 1916 a street in [[the Bronx]], New York City, was named in his honour (Huxley Avenue).<ref>{{cite book |last=McNamara |first=John |author-link= |date=1991 |title=''History in Asphalt'' |url= |location=Harrison, NY |publisher=Harbor Hill Books |page=137 |isbn=0-941980-15-4}}</ref>

As recognition of his many public services, he was appointed [[Privy Councillor]] in 1892.

Despite his many achievements, he was given no award by the British state until late in life. In this, he did better than Darwin, who got no award of any kind from the state. (Darwin's proposed knighthood was vetoed by ecclesiastical advisers, including Wilberforce.)<ref>{{Harvp|Desmond|Moore|1991}}</ref> Perhaps Huxley had commented too often on his dislike of honours, or perhaps his many assaults on the traditional beliefs of organised religion made enemies in the establishment – he had vigorous debates in print with [[Benjamin Disraeli]], [[William Ewart Gladstone]], and [[Arthur Balfour]], and his relationship with [[Lord Salisbury]] was less than tranquil.<ref name=autogenerated4/><ref name=autogenerated1>{{Harvp|Desmond|1997}}</ref>

Huxley was for about thirty years evolution's most effective advocate, and for some Huxley was "''the'' premier advocate of science in the nineteenth century [for] the whole English-speaking world".<ref>{{Harvp|Lyons|1999|p=11}}</ref>

Though he had many admirers and disciples, his retirement and later death left British zoology somewhat bereft of leadership. He had, directly or indirectly, guided the careers and appointments of the next generation. Huxley thought he was "the only man who can carry out my work": The deaths of Balfour and [[William Kingdon Clifford]] were "the greatest losses to science in our time".<ref name=autogenerated4 />

=== Vertebrate palaeontology ===

The first half of Huxley's career as a palaeontologist is marked by a rather strange predilection for 'persistent types', in which he seemed to argue that evolutionary advancement (in the sense of major new groups of animals and plants) was rare or absent in the [[Phanerozoic]]. In the same vein, he tended to push the origin of major groups such as birds and mammals back into the [[Palaeozoic]] era, and to claim that no order of plants had ever gone extinct.<ref>{{Cite web |last=Huxley |first=Thomas Henry |title=Geological Contemporaneity and Persistent Types of Life |url=https://www.gutenberg.org/ebooks/2936/pg2936-images.html |access-date=2024-01-23 |website=Project Gutenberg |language=en}}</ref>

Much paper has been consumed by historians of science ruminating on this strange and somewhat unclear idea.<ref>{{cite book |author=Desmond, A. |year=1982 |title=Archetypes and Ancestors: Palaeontology in Victorian London 1850–1875 |publisher=Muller |place=London, UK}}</ref> Huxley was wrong to pitch the loss of [[order (biology)|orders]] in the Phanerozoic as low as 7%, and he did not estimate the number of new orders which evolved. Persistent types sat rather uncomfortably next to Darwin's more fluid ideas; despite his intelligence, it took Huxley a surprisingly long time to appreciate some of the implications of evolution. However, gradually Huxley moved away from this conservative style of thinking as his understanding of palaeontology, and the discipline itself, developed.{{citation needed|date=May 2017}}

Huxley's detailed anatomical work was, as always, first-rate and productive. His work on [[fossil fish]] shows his distinctive approach: Whereas pre-Darwinian naturalists collected, identified, and classified, Huxley worked mainly to reveal the evolutionary relationships between groups.<ref name=ODNB>{{cite ODNB |title=Huxley, Thomas Henry (1825–1895), biologist and science educationist |url=https://www.oxforddnb.com/view/10.1093/ref:odnb/9780198614128.001.0001/odnb-9780198614128-e-14320 |access-date=20 October 2025 |author=Desmond, Adrian |date=14 September 2023 |url-access=subscription}}</ref>

[[Image:Thomas Henry Huxley by Theodore Blake Wirgman.jpg|left|thumb|Huxley by Wirgman<br/>a drawing in pencil (1882)]]

The [[lobe-finned fish]] (such as [[coelacanth]]s and [[lungfish]]) have paired appendages whose internal skeleton is attached to the shoulder or pelvis by a single bone, the humerus or femur. His interest in these fish brought him close to the origin of [[tetrapod]]s, one of the most important areas of vertebrate palaeontology.<ref>{{Harvp|Clack|2002}}</ref><ref>{{Harvp|Huxley|1861|pp=67–84}}</ref><ref>{{Harvp|Foster|Lankester|2007|pp=163–187}}</ref>

The study of fossil reptiles led to his demonstrating the fundamental affinity of birds and reptiles, which he united under the title of ''[[Sauropsida]]''. His papers on ''[[Archaeopteryx]]'' and the [[origin of birds]] were of great interest then and still are.<ref name=foster1903/><ref>{{Harvp|Paul|2002|pp=171–224}}</ref><ref>{{Harvp|Prum|2003|pp=550–561}}</ref>

Apart from his interest in persuading the world that man was a primate and had descended from the same stock as the apes, Huxley did little work on mammals, with one exception. On his tour of America Huxley was shown the remarkable series of fossil horses, discovered by [[Othniel Charles Marsh]], in [[Yale University]]'s [[Peabody Museum of Natural History|Peabody Museum]].<ref>{{Harvp|Desmond|1997|p=88}}</ref><ref>{{Harvp|Huxley|1877}}</ref> An Easterner, Marsh was America's first professor of palaeontology, but also one who had come west into hostile Indian territory in search of fossils, hunted buffalo, and met [[Red Cloud]] (in 1874).<ref name=Plate-1964>{{cite book |author=Plate, Robert |year=1964 |title=The Dinosaur Hunters: Othniel C. Marsh and Edward D. Cope |publisher=David McKay Company |place=New York, NY}}</ref>{{rp|style=ama|pp= 52, 141–160}} Funded by his uncle [[George Peabody]], Marsh had made some remarkable discoveries: the huge [[Cretaceous]] aquatic bird ''[[Hesperornis]]'', and the dinosaur footprints along the [[Connecticut River]] were worth the trip by themselves, but the horse fossils were really special. After a week with Marsh and his fossils, Huxley wrote excitedly, "The collection of fossils is the most wonderful thing I ever saw."<ref name=Plate-1964/>{{rp|style=ama|p= 203}}

[[File:Eohippus + Eohomo.gif|thumb|Huxley's sketch of then hypothetical five-toed ''[[Eohippus]]'' being ridden by "Eohomo"]]

The collection at that time went from the small four-toed forest-dwelling ''[[Orohippus]]'' from the [[Eocene]] through three-toed species such as ''[[Miohippus]]'' to species more like the modern horse. By looking at their teeth he could see that, as the size grew larger and the toes reduced, the teeth changed from those of a browser to those of a grazer. All such changes could be explained by a general alteration in habitat from forest to grassland.{{citation needed|date=May 2017}} And, it is now known, that is what did happen over large areas of North America from the Eocene to the [[Pleistocene]]: The ultimate causative agent was global temperature reduction (see [[Paleocene–Eocene Thermal Maximum]]). The modern account of the [[evolution of the horse]] has many other members, and the overall appearance of the tree of descent is more like a bush than a straight line.

The horse series also strongly suggested that the process was gradual, and that the origin of the modern horse lay in North America, not in Eurasia. If so, then something must have happened to horses in North America, since none were there when Europeans arrived. The experience with Marsh was enough for Huxley to give credence to Darwin's gradualism, and to introduce the story of the horse into his lecture series.{{Citation needed|date=January 2016}}<ref name=Plate-1964/>{{rp|style=ama|p= 204}}

Marsh's and Huxley's conclusions were initially quite different. However, Marsh carefully showed Huxley his complete sequence of fossils. As Marsh put it, Huxley "then informed me that all this was new to him and that my facts demonstrated the evolution of the horse beyond question, and for the first time indicated the direct line of descent of an existing animal. With the generosity of true greatness, he gave up his own opinions in the face of new truth, and took my conclusions as the basis of his famous New York lecture on the horse."<ref name=Plate-1964/>{{rp|style=ama|p= 204}}

==Support of Darwin==
{{See also|Reactions to On the Origin of Species}}
[[File:Huxley - Mans Place in Nature.jpg|thumb|left|upright=1.3<!--fmt for low img-->|The frontispiece to Huxley's ''[[Evidence as to Man's Place in Nature]]'' (1863) compares ape and human skeletons. The [[gibbon]] (left) is double size.]]

Huxley was originally not persuaded by "development theory", as evolution was once called. This can be seen in his savage review<ref>{{Harvnb|Huxley|1854|}} p.425–439.</ref> of [[Robert Chambers (journalist)|Robert Chambers]]' ''[[Vestiges of the Natural History of Creation]]'', an 1844 book which contained some quite pertinent arguments in favour of evolution. Huxley had also rejected [[Jean-Baptiste Lamarck]]'s theory of transmutation, on the basis that there was insufficient evidence to support it. All this scepticism was brought together in a lecture to the Royal Institution,<ref>{{Harvnb|Huxley|1855|}} p. 82–85.</ref> which made Darwin anxious enough to set about an effort to change young Huxley's mind. It was the kind of thing Darwin did with his closest scientific friends, but he must have had some particular intuition about Huxley, who was by all accounts a most impressive person even as a young man.<ref>{{Harvnb|Browne|1995}}</ref><ref>{{Harvnb|Desmond|1994|}} p. 222.</ref>

Huxley was therefore one of the small group who knew about Darwin's ideas before they were published (the group included [[Joseph Dalton Hooker]] and [[Charles Lyell]]). The first publication by Darwin of his ideas came when Wallace sent Darwin his famous paper on natural selection, which was presented by Lyell and Hooker to the [[Linnean Society]] in 1858 alongside excerpts from Darwin's notebook and a Darwin letter to [[Asa Gray]].<ref name=autogenerated2>{{Harvnb|Browne|2002}}</ref><ref>{{Harvnb|Darwin|Wallace|1858|[http://darwin-online.org.uk/content/frameset?viewtype=text&itemID=F350&pageseq=1 45–62]}}</ref> Huxley's famous response to the idea of natural selection was "How extremely stupid not to have thought of that!"<ref>{{Harvnb|Huxley|1900}} vol. 1, p.189.</ref> However, he never conclusively made up his mind about whether natural selection was the main method for evolution, though he did admit it was a hypothesis which was a good working basis.

Logically speaking, the prior question was whether evolution had taken place at all. It is to this question that much of Darwin's ''[[On the Origin of Species]]'' was devoted. Its publication in 1859 completely convinced Huxley of evolution and it was this and no doubt his admiration of Darwin's way of amassing and using evidence that formed the basis of his support for Darwin in the debates that followed the book's publication.

Huxley's support started with his anonymous favourable review of the ''Origin'' in the ''Times'' for 26 December 1859,<ref>{{Harvnb|Huxley|1893–1894a|pp=1–20}}</ref> and continued with articles in several periodicals, and in a lecture at the [[Royal Institution]] in February 1860.<ref>{{Harvnb|Foster|Lankester|2007|}} p. 400.</ref> At the same time, [[Richard Owen]], whilst writing an extremely hostile anonymous review of the ''Origin'' in the ''[[Edinburgh Review]]'',<ref>{{Harvnb|Owen|1860}}</ref> also primed [[Samuel Wilberforce]] who wrote one in the ''[[Quarterly Review]]'', running to 17,000 words.<ref>{{Harvnb|Wilberforce|1860}}</ref> The authorship of this latter review was not known for sure until Wilberforce's son wrote his biography. So it can be said that, just as Darwin groomed Huxley, so Owen groomed Wilberforce; and both the proxies fought public battles on behalf of their principals as much as themselves. Though we do not know the exact words of the Oxford debate, we do know what Huxley thought of the review in the ''Quarterly'':

[[Image:TH Huxley 41.5 KB.jpg|thumb|Caricature of Huxley by<br />[[Carlo Pellegrini (caricaturist)|Carlo Pellegrini]] in ''[[Vanity Fair (British magazine 1868–1914)|Vanity Fair]]'', 1871]]

<blockquote>Since [[Lord Brougham]] assailed Dr [[Thomas Young (scientist)|Young]], the world has seen no such specimen of the insolence of a shallow pretender to a Master in Science as this remarkable production, in which one of the most exact of observers, most cautious of reasoners, and most candid of expositors, of this or any other age, is held up to scorn as a "flighty" person, who endeavours "to prop up his utterly rotten fabric of guess and speculation," and whose "mode of dealing with nature" is reprobated as "utterly dishonourable to Natural Science."</blockquote>

<blockquote>If I confine my retrospect of the reception of the ''Origin of Species'' to a twelvemonth, or thereabouts, from the time of its publication, I do not recollect anything quite so foolish and unmannerly as the ''Quarterly Review'' article...<ref>Darwin, Francis (ed) 1887. ''The life and letters of Charles Darwin, including an autobiographical chapter''. Murray, London, volume 2.</ref><ref>A more complete version is available in [[q:Thomas Henry Huxley|Wikiquote]]</ref></blockquote>

Since his death, Huxley has become known as "Darwin's Bulldog", taken to refer to his pluck and courage in debate, and to his perceived role in protecting the older man. The sobriquet appears to be Huxley's own invention, although of unknown date,<ref>"'I am Darwin's bull-dog,' he once said ...": {{harvnb|Huxley|1900|loc=Volume 1, p. [https://archive.org/details/dli.granth.90929/page/362/mode/2up 363]}}</ref> and it was not current in his lifetime.<ref name="vanWyheBulldog">{{cite web|url=https://www.linnean.org/news/2019/07/01/1st-july-2019-why-there-was-no-darwins-bulldog|title=Why there was no "Darwin's bulldog"|last=van Wyhe|first=John|author-link=John van Wyhe|date=1 July 2019|publisher=The Linnean Society of London|access-date=12 March 2021|archive-date=15 April 2021|archive-url=https://web.archive.org/web/20210415012354/https://www.linnean.org/news/2019/07/01/1st-july-2019-why-there-was-no-darwins-bulldog|url-status=live}}</ref>

While the second half of Darwin's life was lived mainly within his family, the younger and combative Huxley operated mainly out in the world at large. A letter from Huxley to [[Ernst Haeckel]] (2 November 1871) states: "The dogs have been snapping at [Darwin's] heels too much of late."

===Debate with Wilberforce===
{{Main|1860 Oxford evolution debate}}

Famously, Huxley responded to Wilberforce in the debate at the [[British Association]] meeting, on Saturday 30 June 1860 at the Oxford University Museum. Huxley's presence there had been encouraged on the previous evening when he met Robert Chambers, the Scottish publisher and author of ''Vestiges'', who was walking the streets of Oxford in a dispirited state, and begged for assistance. The debate followed the presentation of a paper by [[John William Draper]], and was chaired by Darwin's former botany tutor [[John Stevens Henslow]]. Darwin's theory was opposed by the Bishop of Oxford, [[Samuel Wilberforce]], and those supporting Darwin included Huxley and their mutual friends Hooker and [[John Lubbock, 1st Baron Avebury|John Lubbock]]. The platform featured [[Sir Benjamin Collins Brodie, 1st Baronet|Sir Benjamin Brodie]] and Professor Beale, and [[Robert FitzRoy]], who had been captain of HMS ''Beagle'' during Darwin's voyage, spoke against Darwin.<ref>Jensen, J. Vernon 1991. ''Thomas Henry Huxley: communicating for science''. U. of Delaware Press, Newark. p. 209, note 67.</ref>

Wilberforce had a track record against evolution as far back as the previous Oxford B.A. meeting in 1847 when he attacked Chambers' ''Vestiges''. For the more challenging task of opposing the ''Origin'', and the implication that man descended from apes, he had been assiduously coached by [[Richard Owen]]{{mdash}}Owen stayed with him the night before the debate.<ref>{{Harvnb|Desmond|Moore|1991}} p. 493</ref> On the day, Wilberforce repeated some of the arguments from his ''Quarterly Review'' article (written but not yet published), then ventured onto slippery ground. His famous jibe at Huxley (as to whether Huxley was descended from an ape on his mother's side or his father's side) was probably unplanned, and certainly unwise. Huxley's reply to the effect that he would rather be descended from an ape than a man who misused his great talents to suppress debate{{mdash}}the exact wording is not certain{{mdash}}was widely recounted in pamphlets and a spoof play.{{citation needed|date = June 2022}}

The letters of [[Alfred Newton]] include one to his brother giving an eyewitness account of the debate, and written less than a month afterwards.<ref>Wollaston AFR 1921. ''[https://archive.org/stream/lifeofalfrednewt01woll#page/118/mode/2up Life of Alfred Newton]: late Professor of Comparative Anatomy, Cambridge University 1866–1907, with a Preface by Sir [[Archibald Geikie]] OM''. Dutton, NY. pp. 118–120.</ref> Other eyewitnesses, with one or two exceptions (Hooker especially thought ''he'' had made the best points), give similar accounts, at varying dates after the event.<ref>Jensen, J. Vernon 1991. ''Thomas Henry Huxley: communicating for science''. U. of Delaware Press, Newark. [Chapter 3 is an excellent survey, and its notes give references to all the eyewitness accounts except Newton: see notes 61, 66, 67, 78, 79, 80, 81, 84, 86, 87, 89, 90, 93, 95: pp. 208–211]</ref> The general view was and still is that Huxley got much the better of the exchange, though Wilberforce himself thought he had done quite well. In the absence of a verbatim report, differing perceptions are difficult to judge fairly; Huxley wrote a detailed account for Darwin, a letter which does not survive; however, a letter to his friend Frederick Daniel Dyster does survive with an account just three months after the event.<ref>Huxley to Dr FD Dyster, 9 September 1860, Huxley Papers 15.117.</ref><ref>{{Harvnb|Browne|2002|}} p. 118.</ref><ref>{{Harvnb|Huxley|1900|}} Chapter 14</ref><ref>
{{Harvnb|Desmond|1994|}} pp. 276–281.</ref><ref name=lucas>{{Harvnb|Lucas|1979|}} p. 313–330. A pro-Wilberforce account; lists many sources, but ''not'' Alfred Newton's letter to his brother. Many of Lucas's points are treated adversely in Jensen 1991, for example, note 77, p. 209.</ref><ref>{{Harvnb|Gould|1991}} Chapter 26 'Knight takes Bishop?' is Gould's take on the Huxley-Wilberforce debate.</ref>

One effect of the debate was to hugely increase Huxley's visibility amongst educated people, through the accounts in newspapers and periodicals. Another consequence was to alert him to the importance of public debate: a lesson he never forgot. A third effect was to serve notice that Darwinian ideas could not be easily dismissed: on the contrary, they would be vigorously defended against orthodox authority.<ref>Darwin F. (ed) 1897–99. ''Life and letters of Charles Darwin''. 2 vols, Murray, London. I, 156-7 Darwin to Huxley: "It is of enormous importance the showing the world that a few first-rate men are not afraid of expressing their opinion."</ref><ref>Darwin F. and A.C. Seward (eds) 1903. ''More letters of Charles Darwin''. 2 vols, Murray, London. II, 204 Leonard Huxley: "The importance... lay in the open resistance that was made to authority".</ref> A fourth effect was to promote professionalism in science, with its implied need for scientific education. A fifth consequence was indirect: as Wilberforce had feared, a defence of evolution did undermine literal belief in the [[Old Testament]], especially the [[Book of Genesis]]. Many of the liberal clergy at the meeting were quite pleased with the outcome of the debate; they were supporters, perhaps, of the controversial ''[[Essays and Reviews]]''. Thus, both on the side of science and on that of religion, the debate was important and its outcome significant.<ref>Jensen, J. Vernon 1991. ''Thomas Henry Huxley: communicating for science''. U. of Delaware Press, Newark. p. 83-86.</ref> (see also [[#Schools and the Bible|below]])

Huxley and Wilberforce remained on courteous terms after the debate (and able to work together on projects such as the Metropolitan Board of Education), whereas Huxley and Owen were never reconciled.{{citation needed|date = June 2022}}<!--rather an opinion, too.-->

===Man's place in nature===
{{See also|Man's Place in Nature}}

For nearly a decade his work was directed mainly to the relationship of man to the apes. This led him directly into a clash with [[Richard Owen]], a man widely disliked for his behaviour whilst also being admired for his capability. The struggle was to culminate in some severe defeats for Owen. Huxley's [[Croonian Lecture]], delivered before the Royal Society in 1858 on ''The Theory of the Vertebrate Skull'' was the start. In this, he rejected Owen's theory that the bones of the skull and the spine were [[homology (biology)|homologous]], an opinion previously held by [[Johann Wolfgang von Goethe]] and [[Lorenz Oken]].<ref>{{Harvnb|Foster|Lankester|2007|}} p. 538–606.</ref>

[[Image:T.H.Huxley 1857.jpg|thumb|Huxley at 32]]

From 1860 to 1863 Huxley developed his ideas, presenting them in lectures to working men, students and the general public, followed by publication. Also in 1862 a series of talks to working men was printed lecture by lecture as pamphlets, later bound up as a little green book; the first copies went on sale in December.<ref>{{Harvnb|Huxley|1862b}}</ref> Other lectures grew into Huxley's most famous work ''Evidence as to Man's place in Nature'' (1863) where he addressed the key issues long before [[Charles Darwin]] published his ''[[Descent of Man]]'' in 1871.{{citation needed|date = June 2022}}

Although Darwin did not publish his ''Descent of Man'' until 1871, the general debate on this topic had started years before (there was even a precursor debate in the 18th century between [[James Burnett, Lord Monboddo]], and [[Georges-Louis Leclerc, Comte de Buffon|Georges-Louis Leclerc]]). Darwin had dropped a hint when, in the conclusion to the ''Origin'', he wrote: "In the distant future... light will be thrown on the origin of man and his history".<ref>{{Harvnb|Darwin|1859|loc= [http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=506 p. 490].}}</ref> Not so distant, as it turned out. A key event had already occurred in 1857 when [[Richard Owen]] presented (to the Linnean Society) his theory that man was marked off from all other mammals by possessing features of the brain peculiar to the genus ''Homo''. Having reached this opinion, Owen separated man from all other mammals in a subclass of its own.<ref>{{Harvnb|Owen|1858|}} p. 1–37.</ref> No other biologist held such an extreme view. Darwin reacted "Man...as distinct from a chimpanzee [as] an ape from a platypus... I cannot swallow that!"<ref>{{Harvnb|Burkhardt|1984}}</ref> Neither could Huxley, who was able to demonstrate that Owen's idea was completely wrong.
[[Image:Huxley7.jpg|thumb|left|Huxley c.1870; sketch is a gorilla skull]]

The subject was raised at the 1860 BA Oxford meeting, when Huxley flatly contradicted Owen, and promised a later demonstration of the facts. In fact, a number of demonstrations were held in London and the provinces. In 1862 at the Cambridge meeting of the B.A. Huxley's friend [[William Henry Flower|William Flower]] gave a public dissection to show that the same structures (the posterior horn of the lateral ventricle and hippocampus minor) were indeed present in apes. The debate was widely publicised, and parodied as the ''[[Great Hippocampus Question]]''. It was seen as one of Owen's greatest blunders, revealing Huxley as not only dangerous in debate but also a better anatomist.{{citation needed|date = June 2022}}

Owen conceded that there was something that could be called a hippocampus minor in the apes, but stated that it was much less developed and that such a presence did not detract from the overall distinction of simple brain size.<ref>{{Harvnb|Cosans|2009}} pp. 109–111</ref>

Huxley's ideas on this topic were summed up in January 1861 in the first issue (new series) of his own journal, the ''Natural History Review'': "the most violent scientific paper he had ever composed".<ref name=autogenerated2 /> This paper was reprinted in 1863 as chapter 2 of ''[[Man's Place in Nature]]'', with an addendum giving his account of the Owen/Huxley controversy about the ape brain.<ref>For the full text of the addendum see [[s:The cerebral structure of man and apes]]</ref> In his ''Collected Essays'' this addendum was removed.{{citation needed|date = June 2022}}

The extended argument on the ape brain, partly in debate and partly in print, backed by dissections and demonstrations, was a landmark in Huxley's career. It was highly important in asserting his dominance of comparative anatomy, and in the long run more influential in establishing evolution amongst biologists than was the debate with Wilberforce. It also marked the start of Owen's decline in the esteem of his fellow biologists.{{citation needed|date = June 2022}}

The following was written by Huxley to [[George Rolleston]] before the BA meeting in 1861:
:"My dear Rolleston... The obstinate reiteration of erroneous assertions can only be nullified by as persistent an appeal to facts; and I greatly regret that my engagements do not permit me to be present at the British Association in order to assist personally at what, I believe, will be ''the seventh public demonstration during the past twelve months'' of the untruth of the three assertions, that the posterior lobe of the cerebrum, the posterior cornu of the lateral ventricle, and the hippocampus minor, are peculiar to man and do not exist in the apes. I shall be obliged if you will read this letter to the Section" Yours faithfully, Thos. H. Huxley.<ref>''Athenaeum'' 21 September 1861, p. 498. [key sentence italicised]</ref>

During those years there was also work on human fossil anatomy and anthropology. In 1862 he examined the [[Neanderthal]] skull-cap, which had been discovered in 1857. It was the first pre-''sapiens'' discovery of a fossil man, and it was immediately clear to him that the brain case was surprisingly large.<ref>{{Harvnb|Huxley|1862a|}}, pp. 420–422</ref> Huxley also started to dabble in [[physical anthropology]], and classified the human [[Race (human categorization)|races]] into nine categories, along with placing them under four general categorisations as Australoid, Negroid, Xanthochroic and Mongoloid. Such classifications depended mainly on physical appearance and certain anatomical characteristics.<ref>Huxley, Thomas Henry. "[http://aleph0.clarku.edu/huxley/SM3/GeoDis.html On the geographical distribution of the chief modifications of mankind] {{Webarchive|url=https://web.archive.org/web/20101212220953/http://aleph0.clarku.edu/huxley/SM3/GeoDis.html |date=12 December 2010 }}". The Journal of the Ethnological Society of London (1869–1870) 2.4 (1870): 404–412.</ref><ref>Brantlinger, Patrick. Dark vanishings: discourse on the extinction of primitive races, 1800–1930. Cornell University Press, 2003. [https://books.google.com/books?id=Ai5XAgAAQBAJ&dq=Amphinesian+huxley&pg=PT153] {{Webarchive|url=https://web.archive.org/web/20230331021833/https://books.google.com/books?id=Ai5XAgAAQBAJ&dq=Amphinesian+huxley&pg=PT153|date=31 March 2023}}</ref>

===Natural selection===
Huxley was certainly not slavish in his dealings with Darwin. As shown in every biography, they had quite different and rather complementary characters. Important also, Darwin was a field naturalist, but Huxley was an anatomist, so there was a difference in their experience of nature. Lastly, Darwin's views on science were different from Huxley's views. For Darwin, natural selection was the best way to explain evolution because it explained a huge range of natural history facts and observations: it solved problems. Huxley, on the other hand, was an empiricist who trusted what he could see, and some things were not easily seen. With this in mind, one can appreciate the debate between them, Darwin writing his letters, Huxley never going quite so far as to say he thought Darwin was right.{{citation needed|date = June 2022}}

Huxley's reservations on natural selection were of the type "until selection and breeding can be seen to give rise to varieties which are infertile with each other, natural selection cannot be proved".<ref>Variously worded in {{Harvnb|Huxley|1860a}}, {{Harvnb|Huxley|1860b}}, {{Harvnb|Huxley|1861}}, {{Harvnb|Huxley|1862b}} and {{Harvnb|Huxley1887}}</ref><ref>{{Harvnb|Poulton|1896}} chapter 18 gives detailed quotations from Huxley and discussion—Darwin's letters to Huxley being not yet published</ref> Huxley's position on selection was agnostic; yet he gave no credence to any other theory. Despite this concern about evidence, Huxley saw that if evolution came about through variation, reproduction and selection then other things would also be subject to the same pressures. This included ideas because they are invented, imitated and selected by humans: ‘The struggle for existence holds as much in the intellectual as in the physical world. A theory is a species of thinking, and its right to exist is coextensive with its power of resisting extinction by its rivals.’<ref>{{Cite journal|last=Huxley|first=T. H.|title=The coming of age of 'The origin of species'|journal=Science|volume=os-1|issue=2|pages=15–17, 20|pmid=17751948|year=1880|doi=10.1126/science.os-1.2.15|s2cid=4061790 }}</ref> This is the same idea as [[meme]] theory put forward by [[Richard Dawkins]] in 1976.<ref>{{Cite book|title=The Selfish Gene|last=Dawkins|first=Richard|publisher=Oxford University Press|year=1976}}</ref>

Darwin's part in the discussion came mostly in letters, as was his wont, along the lines: "The [[empirical evidence]] you call for is both impossible in practical terms, and in any event unnecessary. It's the same as asking to see every step in the transformation (or the splitting) of one species into another. My way so many issues are clarified and problems solved; no other theory does nearly so well".<ref>Letters CD to THH in {{Harvnb|Darwin|Seward|1903}} vol 1, pp. 137–8, 225–6, 230–2, 274, 277, 287</ref>

Huxley's reservation, as Helena Cronin has so aptly remarked, was contagious: "it spread itself for years among all kinds of doubters of Darwinism".<ref>{{harvnb|Cronin|1991|}} p. 397.</ref> One reason for this doubt was that comparative anatomy could address the question of descent, ''but not the question of mechanism''.<ref>{{Harvnb|Mayr|1982}}</ref>

===Pallbearer===
Huxley was a pallbearer at the funeral of [[Charles Darwin]] on 26 April 1882.<ref>[[Doug Linder|Linder, Doug]] (2004) [http://law2.umkc.edu/faculty/projects/ftrials/conlaw/huxleyt.html "Thomas Huxley"] {{Webarchive|url=https://web.archive.org/web/20150224035712/http://law2.umkc.edu/faculty/projects/ftrials/conlaw/huxleyt.html |date=24 February 2015 }}</ref>

==The X Club==
{{Main|X Club}}

In November 1864, Huxley succeeded in launching a dining club, the [[X Club]], composed of like-minded people working to advance the cause of science; not surprisingly, the club consisted of most of his closest friends. There were nine members, who decided at their first meeting that there should be no more. The members were: Huxley, [[John Tyndall]], [[Joseph Dalton Hooker|J. D. Hooker]], [[John Lubbock, 1st Baron Avebury|John Lubbock]] (banker, biologist and neighbour of Darwin), [[Herbert Spencer]] (social philosopher and sub-editor of the Economist), [[William Spottiswoode]] ([[mathematician]] and the Queen's Printer), [[Thomas Archer Hirst|Thomas Hirst]] (Professor of Physics at University College London), [[Edward Frankland]] (the new Professor of Chemistry at the Royal Institution) and [[George Busk]], zoologist and palaeontologist (formerly surgeon for [[HMS Dreadnought (1801)|HMS ''Dreadnought'']]). All except Spencer were [[Fellows of the Royal Society]]. Tyndall was a particularly close friend; for many years they met regularly and discussed issues of the day. On more than one occasion Huxley joined Tyndall in the latter's trips into the Alps and helped with his investigations in [[glaciology]].<ref>{{Harvnb|Huxley|1857|}} p. 241.</ref><ref>{{Harvnb|Tyndall|1896|Ref=CITEREFTyndall1896|}} pp. 338–339, 359, 379–383, 406. "During the summer of 1857, he carefully experimented with coloured liquids on the [[Mer de Glace]] and its tributaries..." ''[[Philosophical Magazine]]'' 1857, vol xiv, p. 241.</ref><ref>{{Harvnb|TyndallHuxley,_Thomas_Henry|1857|}} p. 327–346.</ref>

[[File:Drawing of Thomas Henry Huxley.jpg|thumb|right|From the portrait of A. Legros]]

There were also some quite significant X-Club satellites such as [[William Henry Flower|William Flower]] and [[George Rolleston]], (Huxley protegés), and liberal clergyman [[Arthur Penrhyn Stanley|Arthur Stanley]], the Dean of Westminster. Guests such as [[Charles Darwin]] and [[Hermann von Helmholtz]] were entertained from time to time.<ref>{{Harvnb|Jensen|1970|}} pp. 63–72</ref>

They would dine early on first Thursdays at a hotel, planning what to do; high on the agenda was to change the way the Royal Society Council did business. It was no coincidence that the Council met later that same evening. The first item for the Xs was to get the [[Copley Medal]] for Darwin, which they managed after quite a struggle.

The next step was to acquire a journal to spread their ideas. This was the weekly ''[[The Reader (weekly)|Reader]]'', which they bought, revamped and redirected. Huxley had already become part-owner of the ''[[Natural History Review]]''<ref>{{Harvnb|Desmond|1994|}} pp. 284, 289–290.</ref> bolstered by the support of Lubbock, Rolleston, Busk and Carpenter (X-clubbers and satellites). The journal was switched to pro-Darwinian lines and relaunched in January 1861. After a stream of good articles the ''NHR'' failed after four years; but it had helped at a critical time for the establishment of evolution. The ''Reader'' also failed, despite its broader appeal which included art and literature as well as science. The periodical market was quite crowded at the time, but most probably the critical factor was Huxley's time; he was simply over-committed, and could not afford to hire full-time editors. This occurred often in his life: Huxley took on too many ventures, and was not as astute as Darwin at getting others to do work for him.

However, the experience gained with the ''Reader'' was put to good use when the X Club put their weight behind the founding of ''[[Nature (journal)|Nature]]'' in 1869. This time no mistakes were made: above all, there was a permanent editor (though not full-time), [[Joseph Norman Lockyer|Norman Lockyer]], who served until 1919, a year before his death. In 1925, to celebrate his centenary, ''Nature'' issued a supplement devoted to Huxley.<ref>{{Harvnb|Barr|1997|}} p. 1.</ref>

The peak of the X Club's influence was from 1873 to 1885 as Hooker, Spottiswoode and Huxley were [[List of Presidents of the Royal Society|Presidents of the Royal Society]] in succession. Spencer resigned in 1889 after a dispute with Huxley over state support for science.<ref>{{Harvnb|Desmond|1997|}} p. 191.</ref> After 1892 it was just an excuse for the surviving members to meet. Hooker died in 1911, and Lubbock (now [[Baron Avebury|Lord Avebury]]) was the last surviving member.

Huxley was also an active member of the [[Metaphysical Society]], which ran from 1869 to 1880.<ref>{{Harvnb|Irvine|1955}} Chapter 15</ref> It was formed around a nucleus of clergy and expanded to include all kinds of opinions. Tyndall and Huxley later joined [[The Club (Literary Club)|the Club]] (founded by [[Samuel Johnson|Dr. Johnson]]) when they could be sure that Owen would not turn up.<ref>{{Harvnb|Desmond|1997|}} p. 123.</ref>

==Educational influence==
When Huxley himself was young there were virtually no degrees in British universities in the biological sciences and few courses. Most biologists of his day either were self-taught or took medical degrees. When he retired there were established chairs in biological disciplines in most universities, and a broad consensus on the curricula to be followed. Huxley was the single most influential person in this transformation.

===School of Mines and Zoology===
In the early 1870s, the Royal School of Mines moved to new quarters in South Kensington; ultimately it would become one of the constituent parts of Imperial College London. The move gave Huxley the chance to give more prominence to laboratory work in biology teaching, an idea suggested by practice in German universities.<ref name=autogenerated5 /> In the main, the method was based on the use of carefully chosen types, and depended on the dissection of anatomy, supplemented by microscopy, museum specimens and some elementary physiology at the hands of Foster.

The typical day would start with Huxley lecturing at 9 am, followed by a program of laboratory work supervised by his demonstrators.<ref>{{Harvnb|Osborn|1924}}</ref> Huxley's demonstrators were picked men—all became leaders of biology in Britain in later life, spreading Huxley's ideas as well as their own. [[Michael Foster (physiologist)|Michael Foster]] became Professor of Physiology at Cambridge; [[Ray Lankester]] became Jodrell Professor of Zoology at University College London (1875–91), Professor of Comparative Anatomy at Oxford (1891–98) and Director of the Natural History Museum (1898–1907); S.H. Vines became Professor of Botany at Cambridge; [[William Turner Thiselton-Dyer|W.T. Thiselton-Dyer]] became Hooker's successor at Kew (he was already Hooker's son-in-law!); T. Jeffery Parker became Professor of Zoology and Comparative Anatomy at [[University College, Cardiff]]; and William Rutherford<ref>{{Harvnb|Desmond|1997|}} p. 14, 60.</ref> became the Professor of Physiology at Edinburgh. William Flower, Conservator to the Hunterian Museum, and THH's assistant in many dissections, became Sir [[William Henry Flower|William Flower]], Hunterian Professor of Comparative Anatomy and, later, Director of the Natural History Museum.<ref name=autogenerated1 /> It's a remarkable list of disciples, especially when contrasted with Owen who, in a longer professional life than Huxley, left no disciples at all. Darwin said "No one fact tells so strongly against Owen... as that he has never reared one pupil or follower".<ref>Charles Darwin to Asa Gray 1860 in {{Harvnb|Darwin|Seward|1903|}} p. 153.</ref>

[[Image:Huxley, Thomas Henry (1825 - 1895) by Daniel Downey (1829-1881) - 1863-9.jpg|thumb|left|Photograph of Huxley (c. 1890)]]

Huxley's courses for students were so much narrower than the man himself that many were bewildered by the contrast: "The teaching of zoology by use of selected animal types has come in for much criticism";<ref>{{Harvnb|Lester|1995|}} p. 67.</ref> Looking back in 1914 to his time as a student, Sir [[Arthur Shipley]] said "Darwin's later works all dealt with living organisms, yet our obsession was with the dead, with bodies preserved, and cut into the most refined slices".<ref>{{Harvnb|Wollaston|1921|}} p. 102.</ref> E.W MacBride said "Huxley... would persist in looking at animals as material structures and not as living, active beings; in a word... he was a necrologist.<ref>{{Harvnb|MacBride|1934|}} p. 65.</ref> To put it simply, Huxley preferred to teach what he had actually seen with his own eyes.

This largely [[morphology (biology)|morphological]] program of [[comparative anatomy]] remained at the core of most biological education for a hundred years until the advent of cell and molecular biology and interest in evolutionary ecology forced a fundamental rethink. It is an interesting fact that the methods of the field naturalists who led the way in developing the theory of evolution (Darwin, [[Alfred Russel Wallace]], [[Fritz Müller]], [[Henry Walter Bates|Henry Bates]]) were scarcely represented at all in Huxley's program. Ecological investigation of life in its environment was virtually non-existent, and theory, evolutionary or otherwise, was at a discount. [[Michael Ruse]] finds no mention of evolution or Darwinism in any of the exams set by Huxley, and confirms the lecture content based on two complete sets of lecture notes.<ref>{{Harvnb|Ruse|1997}}</ref>

Since Darwin, Wallace and Bates did not hold teaching posts at any stage of their adult careers (and Műller never returned from Brazil) the imbalance in Huxley's program went uncorrected. It is surely strange that Huxley's courses did not contain an account of the evidence collected by those naturalists of life in the tropics; evidence which they had found so convincing, and which caused their views on evolution by natural selection to be so similar. [[Adrian Desmond]] suggests that "[biology] had to be simple, synthetic and assimilable [because] it was to train teachers and had no other heuristic function".<ref>{{Harvnb|Desmond|1997|}} p. 273, note 20.</ref> That must be part of the reason; indeed it does help to explain the stultifying nature of much school biology. But zoology as taught at all levels became far too much the product of one man.

Huxley was comfortable with comparative anatomy, at which he was the greatest master of the day. He was not an all-round naturalist like Darwin, who had shown clearly enough how to weave together detailed factual information and subtle arguments across the vast web of life. Huxley chose, in his teaching (and to some extent in his research) to take a more straightforward course, concentrating on his personal strengths.

===Schools and the Bible===
Huxley was also a major influence in the direction taken by British schools: in November 1870, he was elected to the [[London School Board]] in its first elections.<ref>{{Harvnb|Desmond|1997|}} p. 19–20.</ref> In primary schooling, he advocated a wide range of disciplines, similar to what is taught today: reading, writing, arithmetic, art, science, music, etc. In secondary education he recommended two years of basic liberal studies followed by two years of some upper-division work, focusing on a more specific area of study. A practical example of the latter is his famous 1868 lecture ''[[On a Piece of Chalk]]'' which was first published as an essay in ''[[Macmillan's Magazine]]'' in London later that year.<ref>{{cite web| url = http://aleph0.clarku.edu/huxley/CE8/Chalk.html| title = On a Piece of Chalk (1868)<!-- Bot generated title -->| access-date = 8 December 2006| archive-date = 29 June 2011| archive-url = https://web.archive.org/web/20110629035513/http://aleph0.clarku.edu/huxley/CE8/Chalk.html| url-status = live}}</ref> The piece reconstructs the geological history of Britain from a simple piece of chalk and demonstrates science as "organized common sense".

Huxley supported the reading of the Bible in schools. This may seem out of step with his agnostic convictions, but he believed that the Bible's significant moral teachings and superb use of language were relevant to English life. "I do not advocate burning your ship to get rid of the cockroaches".<ref>Said of those who wished to abolish all religious teaching, when really all they wanted was to free education from the Church. THH 1873. ''Critiques and Addresses'' p. 90.</ref> However, what Huxley proposed was to create an ''edited version'' of the Bible, shorn of "shortcomings and errors... statements to which men of science absolutely and entirely demur... These tender children [should] not be taught that which you do not yourselves believe".<ref>{{Harvnb|Huxley|1893–1894b|p=397}}</ref><ref>{{Harvnb|Bibby|1959|}} p. 153.</ref> The Board voted against his idea, but it also voted against the idea that public money should be used to support students attending church schools. Vigorous debates took place on such points, and the debates were minuted in detail. Huxley said "I will never be a party to enabling the State to sweep the children of this country into denominational schools".<ref>''School Board Chronicle'' vol. 2, p. 326.</ref><ref name="Bibby 1959 p=155">{{Harvnb|Bibby|1959|}} p. 155.</ref> The Act of Parliament which founded board schools permitted the reading of the Bible but did not permit any denominational doctrine to be taught.

It may be right to see Huxley's life and work as contributing to the secularisation of British society which gradually occurred over the following century. [[Ernst Mayr]] said "It can hardly be doubted that [biology] has helped to undermine traditional beliefs and value systems"<ref>{{Harvnb|Mayr|1982|}} p. 80.</ref>—and Huxley more than anyone else was responsible for this trend in Britain. Some modern [[Christian apologists]] consider Huxley the father of [[antitheism]], though he himself maintained that he was an agnostic, not an atheist. He was, however, a lifelong and determined opponent of almost all [[organised religion]] throughout his life, especially the "[[Catholic Church|Roman Church]] ... carefully calculated for the destruction of all that is highest in the moral nature, in the intellectual freedom, and in the political freedom of mankind".<ref name="Bibby 1959 p=155"/><ref>''School Board Chronicle'' vol 2, p. 360.</ref> In the same line of thought, in an article in ''[[Popular Science]]'', Huxley used the expression "the so-called Christianity of Catholicism", explaining: "I say 'so-called' not by way of offence, but as a protest against the monstrous assumption that Catholic Christianity is explicitly or implicitly contained in any trust-worthy record of the teaching of [[Jesus of Nazareth]]."<ref>Bonnier Corporation. ''[https://books.google.com/books?id=tyoDAAAAMBAJ Popular Science] {{Webarchive|url=https://web.archive.org/web/20230331021838/https://books.google.com/books?id=tyoDAAAAMBAJ |date=31 March 2023 }}'' April 1887, Vol. 30, No. 46. {{ISSN|0161-7370}}. ''Scientific and pseudo-scientific realism'' pp. 789–803</ref>

Originally coining the term in 1869, Huxley elaborated on "[[agnosticism]]" in 1889 to frame the nature of claims in terms of what is knowable and what is not. Huxley states<blockquote>Agnosticism, in fact, is not a creed, but a method, the essence of which lies in the rigorous application of a single principle... the fundamental axiom of modern science... In matters of the intellect, follow your reason as far as it will take you, without regard to any other consideration... In matters of the intellect, do not pretend that conclusions are certain which are not demonstrated or demonstrable.<ref>Huxley, T. H. (Feb, 1889). II. Agnosticism. In Christianity and Agnosticism: A controversy. New York, NY: The Humboldt Publishing Co. Retrieved from https://archive.org/stream/agnosticism00variuoft/agnosticism00variuoft_djvu.txt</ref> </blockquote>Use of that term has continued to the present day (see [[Thomas Henry Huxley and agnosticism]]).<ref>Huxley T. H. 1889. Agnosticism: a rejoinder. In ''Collected Essays vol 5 Science and Christian tradition''. Macmillan, London.</ref> Much of Huxley's agnosticism is influenced by Kantian views on human perception and the ability to rely on rational evidence rather than belief systems.<ref>Lightman, B. (1987) ''The Origins of Agnosticism: Victorian Unbelief and the Limits of Knowledge.''. Baltimore, Johns Hopkins University Press. Retrieved from https://doi.org/10.1086/ahr/94.2.444 {{Webarchive|url=https://web.archive.org/web/20210308014136/https://academic.oup.com/ahr/article/94/2/444/67111 |date=8 March 2021 }}</ref>

In 1893, during preparation for the second [[Romanes Lecture]], Huxley expressed his disappointment at the shortcomings of [[Liberal theology|'liberal' theology]], describing its doctrines as 'popular illusions', and the teachings they replaced 'faulty as they are, appear to me to be vastly nearer the truth'.<ref>{{cite book | last= Huxley | first= Leonard | title= Life and Letters of T H Huxley | volume= 3 | page= 118 | publisher= Wentworth Press | year= 2016 | url= http://www.bookrags.com/ebooks/5799/118.html | archive-url= https://web.archive.org/web/20181004185804/http://www.bookrags.com/ebooks/5799/118.html | archive-date= 4 October 2018 | isbn= 9781371175252 | access-date= 4 October 2018 | url-status= live | df= dmy-all }}</ref>

[[Vladimir Lenin]] remarked that for Huxley "agnosticism serves as a fig-leaf for materialism".<ref>{{cite book|last=Lenin|first=V. I.|title=Materialism and Empirio-criticism|location=Moscow|publisher=Progress Publishers|edition=4|orig-date=1909|date=1964|page=195}}</ref> (See also the [[#Debate with Wilberforce|Debate with Wilberforce]] above.)

===Adult education===
[[File:PXA Vol. 6 p.132 Thomas Henry Huxley.jpg|thumb|left|Thomas Henry Huxley, c. 1885, from a [[carte de visite]]]]
[[File:Huxley - Method and results, 1893 - 4282072 F.jpg|thumb|''Method and results'', 1893]]

Huxley's interest in education went still further than school and university classrooms; he made a great effort to reach interested adults of all kinds: after all, he himself was largely self-educated. There were his lecture courses for working men, many of which were published afterwards, and there was the use he made of journalism, partly to earn money but mostly to reach out to the literate public. For most of his adult life, he wrote for periodicals—the ''[[Westminster Review]]'', the ''Saturday Review'', the ''Reader'', the ''[[Pall Mall Gazette]]'', ''Macmillan's Magazine'', the ''Contemporary Review''. Germany was still ahead in formal science education, but interested people in Victorian Britain could use their initiative and find out what was going on by reading periodicals and using the lending libraries.<ref>{{Harvnb|White|2003|}} p. 69.</ref><ref>Note: articles are listed, and some are available, in The Huxley File at [[Clark University]]</ref>

In 1868 Huxley became Principal of the [[South London Working Men's College]] in [[Blackfriars Road]]. The moving spirit was a portmanteau worker, Wm. Rossiter, who did most of the work; the funds were put up mainly by [[Frederick Denison Maurice|F.D. Maurice]]'s [[Christian Socialists]].<ref>{{Harvnb|Bibby|1959|}} p. 33.</ref><ref>{{Harvnb|Desmond|1994|}} p. 361–362.</ref> At sixpence for a course and a penny for a lecture by Huxley, this was some bargain; and so was the free library organised by the college, an idea which was widely copied. Huxley thought, and said, that the men who attended were as good as any country squire.<ref>{{Harvnb|Desmond|1994}} Chapter 19</ref>

The technique of printing his more popular lectures in periodicals which were sold to the general public was extremely effective. A good example was "The Physical Basis of Life", a lecture given in Edinburgh on 8 November 1868. Its theme—that vital action is nothing more than "the result of the molecular forces of the [[protoplasm]] which displays it"—shocked the audience, though that was nothing compared to the uproar when it was published in the ''[[Fortnightly Review]]'' for February 1869. John Morley, the editor, said "No article that had appeared in any periodical for a generation had caused such a sensation".<ref>{{Harvnb|Morley|1917}} p. 90.</ref> The issue was reprinted seven times and [[protoplasm]] became a household word; ''[[Punch (magazine)|Punch]]'' added 'Professor Protoplasm' to his other soubriquets.

The topic had been stimulated by Huxley seeing the [[cytoplasmic streaming]] in plant cells, which is indeed a sensational sight. For these audiences, Huxley's claim that this activity should not be explained by words such as vitality, but by the working of its constituent chemicals, was surprising and shocking. Today we would perhaps emphasise the extraordinary structural arrangement of those chemicals as the key to understanding what cells do, but little of that was known in the nineteenth century.

When the Archbishop of York thought this 'new philosophy' was based on [[Auguste Comte]]'s [[positivism]], Huxley corrected him: "Comte's philosophy [is just] Catholicism minus Christianity" (Huxley 1893 vol 1 of Collected Essays ''Methods & Results'' 156). A later version was "[positivism is] sheer Popery with M. Comte in the chair of St Peter, and with the names of the saints changed". (lecture on [http://aleph0.clarku.edu/huxley/UnColl/Rdetc/Posit.html ''The scientific aspects of positivism''] Huxley 1870 ''Lay Sermons, Addresses and Reviews'' p.&nbsp;149). Huxley's dismissal of positivism damaged it so severely that Comte's ideas withered in Britain.

==Huxley and the humanities==
[[File:Huxley Collected Essays.png|thumb|Collected essays of Huxley]]
During his life, and especially in the last ten years after retirement, Huxley wrote on many issues relating to the humanities.<ref>{{Harvnb|Barr|1997|}}</ref><ref>Paradis, James G. ''T. H. Huxley: Man's place in nature''. University of Nebraska Press, Lincoln 1978.</ref><ref>Peterson, Houston 1932. ''Huxley: prophet of science''. Longmans Green, London.</ref><ref>Huxley T. H. 1893-4. ''Collected essays'': vol 4 ''Science and Hebrew tradition''; vol 5 ''Science and Christian tradition''; vol 6 ''Hume, with helps to the study of Berkeley''; vol 7 ''Man's place in nature''; vol 9: ''Evolution and ethics, and other essays''. Macmillan, London.</ref>

Perhaps the best known of these topics is ''Evolution and Ethics'', which deals with the question of whether biology has anything particular to say about moral philosophy. Both Huxley and his grandson [[Julian Huxley]] gave [[Romanes Lecture]]s on this theme.<ref>Huxley T.H. and Huxley J. 1947. ''Evolution and ethics 1893–1943''. Pilot, London. In USA as ''Touchstone for ethics'', Harper, N.Y. [includes text from the Romanes lectures of both T. H. Huxley and Julian Huxley]</ref><ref>Paradis, James & Williams, George C 1989. ''Evolution and Ethics: T. H. Huxley's 'Evolution and Ethics', with new essays on its Victorian and sociobiological context''. Princeton, N.J.</ref><ref>Reed J. R. 'Huxley and the question of morality'. In {{Harvnb|Barr|1997|}}</ref> For a start, Huxley dismisses religion as a source of [[moral authority]]. Next, he believes the mental characteristics of man are as much a product of evolution as the physical aspects. Thus, human emotions, intellect, and tendency to prefer living in groups and spending resources on raising young are part and parcel of human evolution and therefore [[Heredity|inherit]]ed.

Despite this, the ''details'' of human values and ethics are not inherited: they are partly determined by human culture, and partly chosen on an individual basis. Morality and duty are often at war with natural instincts; ethics cannot be derived from the ''struggle for existence'': "Of moral purpose I see not a trace in nature. That is an article of exclusively human manufacture."<ref>{{Harvnb|Huxley|1900|}} vol. 2, p. 285.</ref> It is therefore an individual's responsibility to make ethical choices (see [[Ethics]] and [[Evolutionary ethics]]). This seems to put Huxley as a [[Compatibilism|compatibilist]] in the [[Free will|Free Will]] vs [[Determinism]] debate. In this argument, Huxley is diametrically opposed to his long-time friend [[Herbert Spencer]].

Huxley's dissection of [[Jean-Jacques Rousseau]]'s views on man and society is another example of his later work. The essay undermines Rousseau's ideas on man as a preliminary to undermining his ideas on the ownership of property. Characteristic is: "The doctrine that all men are, in any sense, or have been, at any time, free and equal, is an utterly baseless fiction."<ref>Huxley T. H. 1890. The natural inequality of man. ''Nineteenth Century'' January; reprinted in ''Collected Essays'' vol. 1, p. 290–335.</ref>

Huxley's method of argumentation (his strategy and tactics of persuasion in speech and print) is itself much studied.<ref>{{Harvnb|Jensen|1991|}}</ref> His career included controversial debates with scientists, clerics and politicians; persuasive discussions with Royal Commissions and other public bodies; lectures and articles for the general public, and a mass of detailed letter-writing to friends and other correspondents. A large number of textbooks have excerpted his prose for anthologies.<ref>{{Harvnb|Jensen|1991|}}, p. 196.</ref>

==Royal and other commissions==
Huxley worked on ten Royal and other commissions (titles somewhat shortened here).<ref>{{Harvnb|Huxley|1900|}}</ref> The [[Royal Commission]] is the senior investigative forum in the British constitution. A rough analysis shows that five commissions involved science and scientific education; three involved medicine and three involved fisheries. Several involve difficult ethical and legal issues. All deal with possible changes to law and/or administrative practice.

===Royal Commissions===
* 1862: Trawling for herrings on the coast of Scotland.
* 1863–1865: Sea fisheries of the United Kingdom.
* 1870–1871: The [[Contagious Diseases Acts]].
* 1870–1875: Scientific instruction and the advancement of science.
* 1876: The practice of subjugating live animals to scientific experiments ([[vivisection]]).
* 1876–1878: The universities of Scotland.
* 1881–1882: The Medical Acts. [i.e. the legal framework for medicine]
* 1884: Trawl, net and beam trawl fishing.

====Other commissions====
*1866: On the [[Royal College of Science for Ireland]].
*1868: On science and art instruction in Ireland.

== Family ==
{{See also|Huxley family}}
{{stack|
[[File:THH by Collier.jpg|thumb|Pencil drawing of Huxley by his daughter, Marian]]
[[File:Tom&julien-72.jpg|thumb|Huxley with his grandson [[Julian Huxley|Julian]] in 1893]]
[[File:Marian Collier (née Huxley) by John Collier.jpg|thumb|Marian (Mady) Huxley, by her husband [[John Collier (Pre-Raphaelite painter)|John Collier]]]]
}}

In 1855 he married [[Henrietta A. Huxley|Henrietta Anne Heathorn]] (1825–1914),<ref>{{cite news |date=3 June 1914 |title=Death of Mrs Huxley |page=4 |newspaper=The Sun |issue=1227 |location=New South Wales, Australia |url=http://nla.gov.au/nla.news-article224650663 |accessdate=2 March 2023 |via=National Library of Australia |archive-date=31 March 2023 |archive-url=https://web.archive.org/web/20230331021848/https://trove.nla.gov.au/newspaper/article/224650663 |url-status=live }}</ref> an English émigrée whom he had met in [[Sydney, Australia]]. They kept correspondence until he was able to send for her. They had five daughters and three sons:

* Noel Huxley (1856–60), died of [[scarlet fever]].
* Jessie Oriana Huxley (1858<ref>{{cite web|last=Huxley|first=T.H.|title=To Lizzie, March 27, 1858|url=http://aleph0.clarku.edu/huxley/letters/58.html|work=Letters and Diary: 1858|publisher=Clark University|access-date=6 February 2014|archive-date=20 February 2014|archive-url=https://web.archive.org/web/20140220101925/http://aleph0.clarku.edu/huxley/letters/58.html|url-status=live}}</ref> −1927), married architect Fred Waller in 1877.
* [[Marian Huxley]] (1859–87), married artist [[John Collier (Pre-Raphaelite painter)|John Collier]] in 1879.
* [[Leonard Huxley (writer)|Leonard Huxley]], (1860–1933) author, father of [[Julian Huxley|Julian]], [[Aldous Huxley|Aldous]] and [[Andrew Huxley]].
* Rachel Huxley (1862–1934) married civil engineer Alfred Eckersley in 1884; he died in 1895. They were parents of the physicist [[Thomas Eckersley]] and the first BBC Chief Engineer [[Peter Eckersley (engineer)|Peter Eckersley]].
* Henrietta (Nettie) Huxley (1863–1940), married Harold Roller, and travelled Europe as a singer.
* Henry Huxley (1865–1946), became a fashionable general practitioner in [[London]].
* Ethel Huxley (1866–1941), married her sister's widower John Collier in 1889.

Huxley's relationships with his relatives and children were genial by the standards of the day—so long as they lived their lives in an honourable manner, which some did not. After his mother, his eldest sister Lizzie was the most important person in his life until his own marriage. He remained on good terms with his children, more than can be said of many Victorian fathers. This excerpt from a letter to Jessie, his eldest daughter is full of affection:
*"Dearest Jess, You are a badly used young person—you are; and nothing short of that conviction would get a letter out of your still worse used Pater, the bête noir of whose existence is letter-writing. Catch me discussing the Afghan question with you, you little pepper-pot! No, not if I know it..." [goes on nevertheless to give strong opinions of the Afghans, at that time causing plenty of trouble to the [[British Raj]]—see [[Second Anglo-Afghan War]]] "There, you plague—ever your affec. Daddy, THH." (letter 7 December 1878, Huxley L 1900)<ref>{{cite web| url = http://aleph0.clarku.edu/huxley/letters/78.html| title = T. H. Huxley Letters and Diary 1878| access-date = 15 January 2011| archive-date = 8 August 2011| archive-url = https://web.archive.org/web/20110808002124/http://aleph0.clarku.edu/huxley/letters/78.html| url-status = live}}</ref>

Huxley's descendants include children of Leonard Huxley:

* [[Julian Huxley|Sir Julian Huxley]] FRS was the first Director of [[UNESCO]] and a notable evolutionary biologist and humanist.
* [[Aldous Huxley]] was a famous author (''[[Brave New World]]'' 1932, ''[[Eyeless in Gaza (novel)|Eyeless in Gaza]]'' 1936, ''[[The Doors of Perception]]'' 1954).
* [[Andrew Huxley|Sir Andrew Huxley]] OM PRS won the [[Nobel Prize in Physiology or Medicine]] in 1963. He was the second Huxley to become President of the Royal Society.

Other significant descendants of Huxley, such as Sir [[Crispin Tickell]], are treated in the [[Huxley family]].

===Mental problems in the family===
Biographers have sometimes noted the occurrence of mental illness in the Huxley family. His father became "sunk in worse than childish imbecility of mind",<ref>letter THH to eldest sister Lizzie 1853 HP 31.21</ref> and later died in [[Barming Asylum]]; brother George suffered from "extreme mental anxiety"<ref>THH to Lizzie 1858 HP 31.24</ref> and died in 1863 leaving serious debts. Brother James, a well-known psychiatrist and Superintendent of Kent County Asylum, was at 55 "as near mad as any sane man can be".<ref>THH to Lizzie HP 31.44</ref> His favourite daughter, the artistically talented Mady (Marian), who became the first wife of the artist [[John Collier (Pre-Raphaelite painter)|John Collier]], was troubled by mental illness for years. She died of pneumonia in her mid-twenties.<ref>THH to JT 1887 HP 9.164</ref><ref>{{Harvnb|Desmond|1997|}} pp. 175–176</ref>

About Huxley himself, we have a more complete record. As a young apprentice to a medical practitioner, aged thirteen or fourteen, Huxley was taken to watch a post-mortem dissection. Afterwards, he sank into a "deep lethargy" and, though Huxley ascribed this to dissection poisoning, Bibby<ref>{{Harvnb|Bibby|1972|}} p. 7</ref> and others may be right to suspect that emotional shock precipitated the [[clinical depression|depression]]. Huxley recuperated on a farm, looking thin and ill.

The next episode we know of in Huxley's life when he suffered a debilitating depression was on the third voyage of HMS ''Rattlesnake'' in 1848.<ref>{{Harvnb|Huxley|1935}} Chapter 5 'Wanderings of a human soul'</ref> Huxley had further periods of depression at the end of 1871,<ref>{{Harvnb|Desmond|1997|}} p. 27</ref> and again in 1873.<ref>{{Harvnb|Desmond|1997|}} p. 49</ref> Finally, in 1884 he sank into another depression, and this time it precipitated his decision to retire in 1885, at the age of 60.<ref>{{Harvnb|Desmond|1997|}} p. 151</ref> This is enough to indicate the way depression (or perhaps a moderate [[bipolar disorder]]) interfered with his life, yet unlike some of the other family members, he was able to function extremely well at other times.

The problems continued sporadically into the third generation. Two of Leonard's sons suffered serious depression: Trevennen committed suicide in 1914 and Julian suffered a breakdown in 1913,<ref>{{Harvnb|Clark|1968}}</ref> and five more later in life.

== Satires ==
Darwin's ideas and Huxley's controversies gave rise to many cartoons and satires. It was the debate about man's place in nature that roused such widespread comment: cartoons are so numerous as to be almost impossible to count; Darwin's head on a monkey's body is one of the visual clichés of the age. The "[[Great Hippocampus Question]]" attracted particular attention:

* "Monkeyana"<ref>{{cite magazine |author=Egerton, P. (pseudonym='Gorilla') |author-link=Sir Philip Grey Egerton, 10th Baronet |date=18 May 1861 |title=Monkeyana |magazine=Punch |volume=40}}</ref> signed 'Gorilla'. this turned out to be by [[Sir Philip Grey Egerton, 10th Baronet|Sir Philip Egerton MP]], amateur naturalist, fossil fish collector, and Richard Owen's patron.<ref>{{Harvp|Desmond|1994|p=296}}</ref> The last two stanzas include a reference to Huxley's comment that "Life is too short to occupy oneself with the slaying of the slain more than once.":<ref>{{cite book |title=The Athenaeum |year=1861 |publisher=British Periodicals Limited |page=498 |url=https://books.google.com/books?id=evbfIPxyEW4C&pg=PA498}}</ref> <blockquote>Next HUXLEY replies<br/>That OWEN he lies<br/>And garbles his Latin quotation;<br/>That his facts are not new,<br />His mistakes not a few,<br/>Detrimental to his reputation.<br/><br/>To twice slay the slain<br/>By dint of the Brain<br/>(Thus HUXLEY concludes his review)<br/>Is but labour in vain,<br/>unproductive of gain,<br />And so I shall bid you "Adieu"!</blockquote>
* "The Gorilla's dilemma":<ref>{{cite magazine |title=The Gorilla's dilemma |magazine=Punch |year=1862 |volume=43 |page=164}}</ref> First two lines: <blockquote>Say am I a man or a brother,<br/>Or only an anthropoid ape?</blockquote>
* ''Report of a sad case recently tried before the Lord Mayor, Owen versus Huxley''.<ref>{{cite book |title=Pamphlet |publisher=George Pycraft |place=London, UK |year=1863}} cited in {{cite book |title=Huxley Papers |at=79.6}}</ref> Lord Mayor asks whether either side is known to the police: <blockquote>Policeman&nbsp;X — Huxley, your Worship, I take to be a young hand, but very vicious; but Owen I have seen before. He got into trouble with an old bone man, called Mantell, who never could be off complaining as Owen prigged his bones. People did say that the old man never got over it, and Owen worritted him to death; but I don't think it was so bad as that. Hears as Owen takes the chair at a crib in Bloomsbury. I don't think it will be a harmonic meeting altogether. And Huxley hangs out in Jermyn Street.</blockquote>
Huxley's low set included Hooker "in the green and vegetable line" and "Charlie Darwin, the pigeon-fancier"; Owen's "crib in Bloomsbury" was the British Museum, of which Natural History was but one department. [[Jermyn Street]] is known for its shops of men's clothing, possibly implying that Huxley was a [[dandy]].

[[File:Water-babies 2.jpg|thumb|Huxley (right) and Richard Owen inspect a "water baby" in [[Edward Linley Sambourne]]'s illustration (1881).]]

* ''[[The Water-Babies, A Fairy Tale for a Land Baby|The Water Babies: A fairy tale for a land baby]]'' by [[Charles Kingsley]] (serialised in ''Macmillan's Magazine'' 1862–1863, published in book form, with additions, in 1863). Kingsley had been among first to give a favourable review to Darwin's ''[[On the Origin of Species]]'', having "long since ... learnt to disbelieve the dogma of the permanence of species",<ref>{{Harvnb|Darwin|1887|p=[http://darwin-online.org.uk/content/frameset?itemID=F1452.2&viewtype=text&pageseq=303 287]}}</ref> and the story includes a [[satire]] on the [[reaction to Darwin's theory]], with the main scientific participants appearing, including [[Richard Owen]] and Huxley. In 1892, Huxley's five-year-old grandson [[Julian Huxley|Julian]] saw the illustration by [[Edward Linley Sambourne|Sambourne]] (right) and wrote his grandfather a letter asking: <blockquote>Dear Grandpater — Have you seen a Waterbaby? Did you put it in a bottle? Did it wonder if it could get out? Could I see it some day? — Your loving Julian.</blockquote> Huxley wrote back: <blockquote>My dear Julian — I could never make sure about that Water Baby ... My friend who wrote the story of the Water Baby was a very kind man and very clever. Perhaps he thought I could see as much in the water as he did — There are some people who see a great deal, and some who see very little in the same things.<br/><br/>When you grow up I dare say you will be one of the great-deal seers, and see things more wonderful than the Water Babies where other folks can see nothing.</blockquote>

== Racism debate ==
In October 2021 a history group reviewing colonial links told [[Imperial College London]] that, because Huxley "might now be called racist", it should remove a bust of him and rename its Huxley Building. The group of 21 academics had been launched in the wake of [[Black Lives Matter]] protests in 2020 to address Imperial's "links to the [[British Empire]]" and build a "fully inclusive organisation". It reported that Huxley's paper ''Emancipation – Black and White'' "espouses a racial hierarchy of intelligence" that helped to feed ideas around [[eugenics]], which "falls far short of Imperial's modern values", and that his theories "might now be called 'racist' in as much as he used racial divisions and hierarchical categorisation in his attempt to understand their origins in his studies of human evolution". Imperial's provost [[Ian Walmsley]] responded that Imperial would "confront, not cover up, uncomfortable or awkward aspects of our past" and this was "very much not a '[[cancel culture]]' approach". Students would be consulted before management decided on any action to be taken.<ref>{{cite news|url=https://www.telegraph.co.uk/news/2021/10/26/imperial-college-told-remove-bust-slavery-abolitionist-might/|archive-url=https://ghostarchive.org/archive/20220112/https://www.telegraph.co.uk/news/2021/10/26/imperial-college-told-remove-bust-slavery-abolitionist-might/|archive-date=12 January 2022|url-access=subscription |url-status=live|last=Somerville|first=Ewan|title=Imperial College told to remove bust of slavery abolitionist because he 'might now be called racist'|work=The Telegraph|location=London|date=26 October 2021|access-date=28 October 2021}}{{cbignore}}.</ref>

A critical response by [[Nick Matzke]] said that Huxley was a public, longstanding [[abolitionist]] who wrote "the most complete demonstration of the specific diversity of the types of mankind will nowise constrain science to spread her ægis over their [slaveholders'] atrocities" and "the North is justified in any expenditure of blood or of money, which shall eradicate a system hopelessly inconsistent with the moral elevation, the political freedom, or the economical progress of the American people";<ref>{{cite web|url=https://mathcs.clarku.edu/huxley/comm/Books/Taylor.html|last=Huxley|first=Thomas Henry|title=Professor Huxley on the Negro Question|website=Ladies London Emancipation Society 1864|date=1864|access-date=18 November 2021|via=Clark University|archive-date=10 August 2022|archive-url=https://web.archive.org/web/20220810162246/https://mathcs.clarku.edu/huxley/comm/Books/Taylor.html|url-status=live}}</ref> a major opponent of the racist position of [[polygenism]], as well as the position that some human races were transitional (in 1867 Huxley said "there was no shade of justification for the assertion that any existing modification of mankind now known was to be considered as an intermediate form between man and the animals next below him in the scale of the fauna of the world"); a vehement opponent of the scientific racist [[James Hunt (speech therapist)|James Hunt]]; and a political radical who believed in granting equal rights and the vote to both black people and women.<ref>{{cite web|url=http://pandasthumb.org/archives/2021/11/updated-creationists-social-justice-advocates-unite-take-down-huxley.html|last=Matzke|first=Nicholas J.|title=Creationists and advocates of social justice unite to take down T.H. Huxley, a leader in educational inclusion|website=The Panda's Thumb|date=3 November 2021|access-date=18 November 2021|archive-date=14 November 2021|archive-url=https://web.archive.org/web/20211114205237/http://pandasthumb.org/archives/2021/11/updated-creationists-social-justice-advocates-unite-take-down-huxley.html|url-status=live}}</ref>

==Cultural references==
[[File:THOMAS HENRY HUXLEY 1825-1895 Biologist Lived Here.jpg|thumb|[[Blue plaque]] commemorating Huxley in [[Marlborough Place]] in London]]
* Huxley appears alongside Charles Darwin and Samuel Wilberforce in the 2003 play ''[[Darwin in Malibu]]'', written by [[Crispin Whittell]], and is portrayed by [[Toby Jones]] in the 2009 film ''[[Creation (2009 film)|Creation]]''.
* Huxley is referred to as the tutor of the main character, Edward Prendick, in [[H. G. Wells]]' 1896 science fiction novel ''[[The Island of Dr Moreau]]''.
* In ''[[Horse Feathers]]'', a 1932 [[Marx Brothers]] film, [[Groucho Marx]] portrays Prof. Quincy Adams Wagstaff, Dean of Huxley College, while its rival team is Darwin College.


==See also==
* [[Epiphenomenalism]]
* [[European and American voyages of scientific exploration]]
* [[List of presidents of the Royal Society]]
* [[Huxley Memorial Medal and Lecture]]
* [[The Huxley Lecture]]

==References==
{{Reflist|30em}}

==Sources==
{{refbegin|20em}}
* {{citation | last = Encyclopædia Britannica Online | author-link = Encyclopædia Britannica | year = 2006 | title = Thomas Henry Huxley | publisher = [[Encyclopædia Britannica Inc.]] | url = http://www.britannica.com/EBchecked/topic/277746/T-H-Huxley | access-date = 16 September 2009 | archive-date = 26 November 2010 | archive-url = https://web.archive.org/web/20101126163723/http://www.britannica.com/EBchecked/topic/277746/T-H-Huxley | url-status = live }}
* {{citation | editor-last = Barr | editor-first = Alan P. | year = 1997 | title = Thomas Henry Huxley's Place in Science and Letters: Centenary Essays |  location = Georgia | publisher = Athens 
}}
* {{citation | last = Bibby | first = Cyril | author-link = Cyril Bibby |title = T.H. Huxley: scientist, humanist and educator | publisher = Watts | year = 1959 | location = London }}
* {{citation | last = Bibby | first = Cyril | author-link = Cyril Bibby |title = Scientist Extraordinary: the life and work of Thomas Henry Huxley 1825–1895 | publisher = Pergamon | year = 1972 | location = Oxford }}
* {{citation | last = Browne | first = Janet | author-link = Janet Browne| title = Charles Darwin| volume= 1: Voyaging | publisher = Cambridge University Press | year = 1995 }}
* {{citation | last = Browne | first = Janet | title = Charles Darwin| volume= 2: The Power of Place | publisher = Cambridge University Press | year = 2002 }}
* {{citation | editor-last = Burkhardt | editor-first = F.|display-editors= etal | title = The Correspondence of Charles Darwin | publisher = Cambridge University Press | date = 1984|orig-date=1984 onwards: continuing series }}
* {{citation | last = Clack | first = Jenny | title = Gaining Ground: The Origin of Tetrapods |  year = 2002 | location = Indiana }}
* {{citation | last = Clark | first = Ronald W. | author-link = Ronald W. Clark| title = The Huxleys |  year = 1968 | location = London |publisher = Heinemann}}
* {{citation | last = Cosans | first = Christopher | title = Owen's Ape and Darwin's Bulldog: Beyond Darwinism and Creationism | publisher = Indiana University Press | year = 2009 | location = Bloomington }}
* {{citation | last = Cronin | first = Helena | title = The Ant and the Peacock: Altruism and Sexual Selection from Darwin to Today | publisher = Cambridge University Press | year = 1991 }}
* {{citation | last = Darwin | first = Charles | author-link = Charles Darwin | year = 1887 | editor-last = Darwin | editor-first = Francis | editor-link = Francis Darwin | title = The life and letters of Charles Darwin, including an autobiographical chapter | location = London | publisher = John Murray | volume = 2 | url = http://darwin-online.org.uk/content/frameset?itemID=F1452.2&viewtype=text&pageseq=1 | access-date = 23 July 2007 | archive-date = 1 August 2011 | archive-url = https://web.archive.org/web/20110801014542/http://darwin-online.org.uk/content/frameset?itemID=F1452.2&viewtype=text&pageseq=1 | url-status = live }}
* {{citation | last1 = Darwin | first1 = Charles | author-link = Charles Darwin | last2 = Wallace | first2 = Alfred Russel | author2-link = Alfred Russel Wallace | date = 1858 | title = On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection | periodical = Journal of the Proceedings of the Linnean Society of London. Zoology | place = London | volume = 3 | issue = 9 | pages = (Read 1 July): 45–62 | doi=10.1111/j.1096-3642.1858.tb02500.x
| title-link = On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection | doi-access = free }}
*{{citation | last = Darwin | first = Charles | year = 1859 | title = On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life | edition = 1st | location = London | publisher = John Murray | url = http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=1 | access-date = 1 March 2011 | archive-date = 5 October 2008 | archive-url = https://web.archive.org/web/20081005185317/http://darwin-online.org.uk/content/frameset?itemID=F373&viewtype=text&pageseq=1 | url-status = live }}
* {{citation | last1 = Darwin | first1 = Francis | author-link = Francis Darwin | year = 1903 | last2 = Seward | first2 = A.C. | title = More Letters of Charles Darwin. 2 vols | location = London | publisher = John Murray 
}}
* {{citation | last = Desmond | first = Adrian | author-link = Adrian Desmond | year = 1994 | title = Huxley: vol 1 The Devil's Disciple | location = London | publisher = Michael Joseph | isbn = 0-7181-3641-1 }}
* {{citation | last = Desmond | first = Adrian | title = Huxley: vol 2 Evolution's High Priest | publisher = Michael Joseph | year = 1997 | location = London }}
* {{citation | last = Desmond | first = Adrian | title = Huxley: vol 1 and 2 | publisher = Penguin | year = 1998 | location = London }}
* {{citation | last1 = Desmond | first1 = Adrian | year = 1991 | last2 = Moore | first2 = James | title = Darwin | location = London | publisher = Penguin Books, Ltd. 
}}
* {{citation | last = Di Gregorio | first = Mario A | title = T.H. Huxley's Place in Natural Science | publisher = Yale University Press | year = 1984 | location = New Haven | isbn = 0-300-03062-2 | url-access = registration | url = https://archive.org/details/thhuxleysplacein0000digr }}
* {{citation | last = Duncan | first = David | title = Life and Letters of Herbert Spencer. 2 vols | publisher = Michael Joseph | year = 1908 }}
* {{citation | last1 = Eve | first1 = A.S. | year = 1945 | last2 = Creasey | first2 = C.H. | title = Life and work of John Tyndall | location = London | publisher = Macmillan | pages = 189–190 | volume = 156 | issue = 3955 | journal = Nature|  doi = 10.1038/156189a0 | bibcode = 1945Natur.156..189R
| s2cid = 4031321 }}
* {{citation | last1 = Foster | first1 = Michael | author-link1 = Michael Foster (physiologist) | last2 = Lankester | first2 = E. Ray | author-link2 = Ray Lankester | title = The scientific memoirs of Thomas Henry Huxley. 4 vols and supplement | publisher = Macmillan | publication-date = 1898–1903 | location = London | isbn = 978-1-4326-4011-8 | year = 2007 }}
* {{citation | last = Galton | first = Francis | title = Hereditary Genius ''(2nd ed.)'' |  year = 1892 | location = London | pages = xix }}
* {{citation | last = Gould | first = Stephen Jay | author-link = Stephen Jay Gould| title = Bully for Brontosaurus | publisher = Random House | year = 1991 }}
* {{citation | first = Linda Z | last = Holland |  title = A chordate with a difference | journal = [[Nature (journal)|Nature]] | issue = 447/7141, pp. 153–155 | publisher = Nature Publishing Group | year = 2007 | location = UK | issn = 0028-0836 | doi = 10.1038/447153a | volume = 447 | pages = 153–5 | pmid = 17495912 |bibcode = 2007Natur.447..153H | s2cid = 5549210 | doi-access = free }}
* {{citation | last = Huxley | first = Julian | author-link = Julian Huxley | title = T.H. Huxley's Diary of the Voyage of HMS Rattlesnake | journal = Nature | volume = 137 | issue = 3454 | pages = 48–49 | publisher = Chatto & Windus | year = 1935 | location = London | bibcode = 1936Natur.137...48Y | doi = 10.1038/137048a0 | s2cid = 41532921 }}
* {{citation | last = Huxley | first = Leonard | author-link = Leonard Huxley (writer) | title = Life and Letters of Thomas Henry Huxley. 2 vols 8vo | publisher = Macmillan | year = 1900 | location = London }}
* {{citation | last = Huxley | first = Thomas Henry | title = Review of Vestiges of the Natural History of Creation, tenth edition | journal = British and Foreign Medico-Chirurgical Review |  issue = 13 |  year = 1854 }}
* {{citation | last = Huxley | first = Thomas Henry | title = On certain zoological arguments commonly adduced in favour of the hypothesis of the progressive development of animal life in time | journal = Proceedings of the Royal Institution 2 (1854–58) | year = 1855 }}
* {{citation | last = Huxley | first = Thomas Henry | author-link = <!--Thomas Henry Huxley --> | title = untitled letter on theory of glaciers | journal = [[Philosophical Magazine]] | volume = xiv | page = 241 | year = 1857}}
* {{citation | last = Huxley | first = Thomas Henry | author-link = <!--Thomas Huxley--> | title = The Oceanic Hydrozoa | publisher = The Ray Society | year=1984 | orig-year=1859 | location = London | isbn = 0-300-03062-2 | url-access = registration | url = https://archive.org/details/thhuxleysplacein0000digr }}
* {{citation | last = Huxley | first = Thomas Henry | title = On species, and races and their origin | journal = Proc. Roy. Inst. 1858–62 |  issue = III |  year = 1860a | page = 195 }}
* {{citation | last = Huxley | first = Thomas Henry | title = The origin of species | journal = Westminster Review |  issue = April |  year = 1860b }}
* {{cite wikisource | last1=Huxley | first1=Thomas Henry | title=On the zoological relations of man with the lower animals | wslink=Natural History Review/Series 2/Volume 1/Number 1/On the Zoological Relations of Man with the Lower Animals | work=Natural History Review | series=2 | volume=1 | number=1 | year=1861 | pages=67–84 | firsticon=yes | noicon=yes }}
* {{citation | first = Thomas Henry | last = Huxley | title = On the fossil remains of Man | journal = Proceedings of the Royal Institution (1858–62) | volume = III |  publisher = The Royal Institution | year = 1862a | location = London }}
* {{citation | last = Huxley | first = Thomas Henry | title = On our knowledge of the causes of the phenomena of organic nature |  year = 1862b | location = London 
}}
* {{citation | last = Huxley | first = Thomas Henry | title = Evidence as to Man's Place in Nature | publisher = Williams & Norwood | year = 1863 | location = London 
}}
* {{citation | last = Huxley | first = Thomas Henry | title = Further remarks on the human remains from the Neanderthal | journal = Natural History Review | issue = 4 | year = 1864 | location = London | pages = 429–46 
}}
* {{citation | last = Huxley | first = Thomas Henry | title = Lay Sermons, Addresses and Reviews | year = 1870 | location = London | pages = 22–23 | issue = 54 | volume = 3 | journal = Nature | doi = 10.1038/003022a0 | bibcode = 1870Natur...3...22G | s2cid = 4071308 | url = https://zenodo.org/record/1429187 | access-date = 6 July 2019 | archive-date = 22 March 2020 | archive-url = https://web.archive.org/web/20200322191847/https://zenodo.org/record/1429187 | url-status = live }}
* {{citation | last = Huxley | first = Thomas Henry | title = American Addresses. |  year = 1877 }}
* {{citation | last = Huxley | first = Thomas Henry | year = 1887 | contribution = On the reception of the 'Origin of Species' | editor-last = Darwin | editor-first = Francis | title = Life & Letters of Charles Darwin | location = London | publisher = John Murray
}}
* {{citation | last = Huxley | first = Thomas Henry | title = Collected essays. 9 vols. Vol 1: Methods and results; vol 2: Darwiniana; vol 3: Science and education; vol 4: Science and Hebrew tradition; vol 5: Science and Christian tradition; vol 6 :Hume, with helps to the study of Berkeley; vol 7:Man's place in nature; vol 8: Discourses biological and geological; vol 9: Evolution and ethics, and other essays | publisher = Macmillan | date = 1893{{ndash}}94 | location = London }}
* {{citation | last = Huxley | first = Thomas Henry | title = Collected essays: vol 2 Darwiniana | publisher = Macmillan | date = 1893–1894a | location = London }}
* {{citation | last = Huxley | first = Thomas Henry | title = Collected essays: vol 3 Science and education | publisher = Macmillan | date = 1893–1894b | location = London }}
* {{citation | last = Huxley | first = Thomas Henry | title = Darwiniana: Essays | publisher = D. Appleton and Company | date = 1898 | location = New York }} Not to be confused with ''[[Darwiniana]]: Essays and Reviews Pertaining to Darwinism'', by [[Asa Gray]].
* {{citation | last = Huxley | first = Thomas Henry | publication-date = 1898–1903 | contribution = Preliminary essay upon the systematic arrangement of the fishes of the Devonian epoch | editor-last = Foster | editor-first = Michael | editor2-last = Lankester | editor2-first = E. Ray | title = The scientific memoirs of Thomas Henry Huxley| volume= 2 | publisher = Macmillan | location = London | pages = 421–460 | isbn = 978-1-4326-4011-8 | year = 2007 }}
* {{cite book|last=Irvine|first=William|title=Apes, Angels and Victorians|year=1955|location=New York}}
* {{citation | last = Jensen | first = J. Vernon | title = The X Club: Fraternity of Victorian Scientists | journal = British Journal for the History of Science | volume = 5 | issue = 1 | pages = 63–72 |  year = 1970 |  doi = 10.1017/S0007087400010621 | pmid = 11609564 | s2cid = 29405737 }}
* {{citation | last = Jensen | first = J. Vernon | title = Thomas Henry Huxley: Communicating for Science. |  publisher = University of Delaware | year = 1991 | location = Newark }}
* {{citation | last = Lester | first = Joe | title = E. Ray Lankester: The Making of Modern British Biology ''(edited, with additions, by Peter J. Bowler)'' | publisher = BSHS Monograph #9 | year = 1995 }}
* {{citation | last = Lucas | first = John R. | author-link = John Lucas (philosopher) | title = Wilberforce and Huxley: A Legendary Encounter | journal = The Historical Journal | volume = 22 | issue = 2 | publisher = [[Cambridge University Press]] | year = 1979 | url = http://users.ox.ac.uk/~jrlucas/legend.html | doi = 10.1017/S0018246X00016848 | pmid = 11617072 | access-date = 9 June 2007 | pages = 313–30 | s2cid = 19198585 | archive-date = 27 June 2020 | archive-url = https://web.archive.org/web/20200627072321/http://users.ox.ac.uk/~jrlucas/legend.html | url-status = dead | url-access = subscription }}
* {{citation | last = Lyons | first = Sherrie L. | title = Thomas Henry Huxley: The Evolution of a Scientist |  year = 1999 | location = New York }}
* {{citation | last = MacBride | first = E.W. | title = Huxley | publisher = Duckworth | year = 1934 | location = London }}
* {{citation | last = MacGillivray | first = John | author-link = John MacGillivray | title = Narrative of the Voyage of HMS Rattlesnake. 2 vols | publisher = Boone | year = 1852 | location = London }}
* {{citation | year = 1982 | editor-last = Mackenzie | editor-first = Norman | editor2-last = Mackenzie | editor2-first = Jenne | title = The Diaries of Beatrice Webb, vol. 1, 1873–1892 | location = London | publisher = Virago
}}
* {{citation | last = Mayr | first = Ernst | author-link = Ernst Mayr| title = The Growth of Biological Thought | publisher = Harvard University Press | year = 1982 }}
* {{citation | last1 = McMillan | first1 = N.D. | last2 = Meehan | first2 = J | title = John Tyndall: 'X'emplar of Scientific & Technological Education | publisher = National Council for Educational Awards | year = 1980 | location = Dublin | url = https://books.google.com/books?id=TPaIAAAACAAJ&q=John+Tyndall:+%27X%27emplar+of+scientific+%26+technological+education | access-date = 14 February 2014 | isbn = 9780905717098 | archive-date = 3 November 2022 | archive-url = https://web.archive.org/web/20221103120253/https://books.google.com/books?id=TPaIAAAACAAJ&q=John+Tyndall:+%27X%27emplar+of+scientific+%26+technological+education | url-status = live }}. (despite its chaotic organisation, this little book contains some nuggets that are well worth sifting)
* {{citation | last = Morley | first = John | title = Recollections. 2 vols | publisher = Macmillan | year = 1917 }}
* {{citation | last = Osborn | first = Henry Fairfield | title = Impressions of Great Naturalists |  year = 1924 }}
* {{citation | last = Owen | first = Richard | author-link = Richard Owen |title = On the characters, principles of division, and primary groups of the Class Mammalia | journal = Proc Linnean Society: Zoology |  issue = 2 | pages = 1–37 |  date = 1858 }}
* {{citation | last = Owen | first = Richard | author-link = Richard Owen |title = Darwin on the Origin of Species | journal = Edinburgh Review |  issue = 111 | pages = 487–532 |  date = 1860 }}
* {{citation | last1 = Paradis | first1 = James | last2 = Williams | first2 = George C.| title = Evolution and Ethics: T. H. Huxley's 'Evolution and Ethics', with New Essays on Its Victorian and Sociobiological Context | publisher = Princeton University Press | year = 1989 | location = Princeton, N.J }}
* {{citation | last = Paradis | first = James G. | title = T.H. Huxley: Man's Place in Nature | publisher = University of Nebraska Press, Lincoln | year = 1978 }}
* {{citation | last = Paul | first = G. | title = Dinosaurs of the Air: The Evolution and Loss of Flight in Dinosaurs and Birds | pages = [https://archive.org/details/dinosaursofairev0000paul/page/171 171–224] | publisher = Johns Hopkins University Press | location = Baltimore | year = 2002 | isbn = 0-8018-6763-0 | url = https://archive.org/details/dinosaursofairev0000paul/page/171 }}
* {{citation | last = Peterson | first = Houston | title = Huxley: Prophet of Science | publisher = Longmans, Green | year = 1932 | location = London }}.
* {{citation | last = Poulton | first = Edward Bagnall | author-link = Edward Bagnall Poulton |title = Charles Darwin and the Theory of Natural Selection | publisher = Cassell | year = 1896 | location = London }}.(Chapter 18 deals with Huxley and natural selection)
* {{citation | last = Pritchard | first = M. | title = A directory of London photographers 1891–1908 |  year = 1994 }}
* {{citation | last = Prum | first = R | title = Are current critiques of the theropod origin of birds science? Rebuttal To Feduccia 2002 | journal = [[The Auk]] | volume = 120 | issue = 2 | pages = 550–561 |  year = 2003 |  doi = 10.1642/0004-8038(2003)120[0550:ACCOTT]2.0.CO;2 | doi-broken-date = 1 July 2025 | s2cid = 85696237 | doi-access =  }}
* {{citation | last = Ruse | first = Michael | author-link = Michael Ruse | year = 1997 | contribution = Thomas Henry Huxley and the Status of Evolution as Science | editor-last = Barr | editor-first = Alan P. | title = Thomas Henry Huxley's place in science and letters: centenary essays | location = Georgia | publisher = Athens
}}
* {{citation | last = Spencer | first = Herbert | author-link = Herbert Spencer| title = Autobiography. 2 vols | publisher = Williams & Norgate | year = 1904 | location = London }}
* {{citation | last1 = Tyndall | first1 = John | author-link = John Tyndall | author2 = Huxley, Thomas Henry | title = On the Structure and Motion of Glaciers | journal = [[Philosophical Transactions]] | volume = 147 | pages = 327–346 | year = 1857 | doi = 10.1098/rstl.1857.0016 | bibcode = 1857RSPT..147..327T | url = https://zenodo.org/record/1432424 | doi-access = free | access-date = 11 September 2019 | archive-date = 22 March 2020 | archive-url = https://web.archive.org/web/20200322191850/https://zenodo.org/record/1432424 | url-status = live }}
* {{citation | title = The Glaciers of the Alps | last = Tyndall | first = John | author-link = John Tyndall | year = 1896 | publisher = [[Longman|Longmans, Green and Co.]] | url = https://archive.org/details/glaciersalpsbeing00tyndrich | edition = Original edition 1860 }}
* {{citation | last = Webb | first = Beatrice | title = My Apprenticeship | journal = Nature | publisher = Longmans | year = 1926 | volume = 118 | issue = 2980 | page = 831 | location = London | doi = 10.1038/118831a0 | bibcode = 1926Natur.118..831C }}
* {{citation | last = Wilberforce | first = Samuel | author-link = Samuel Wilberforce |title = Darwin's Origin of Species | journal = Quarterly Review |  issue = 102 | pages = 225–64 |  date = 1860 }}
* {{citation | last = Wollaston | first = A.F.R. | title = Life of Alfred Newton 1829–1907 |  year = 1921 }}
* {{citation | last = White | first = Paul | title = Thomas Huxley: Making the 'Man of Science' | publisher = Cambridge University Press | year = 2003 }}
{{refend}}

==Selected works==
===Books===

* ''On Our Knowledge of the Causes of the Phenomena of Organic Nature'' (1862)
* ''Evidence as to Man’s Place in Nature'' (1863)
* ''Lay Sermons, Addresses and Reviews'' (1870)
* ''Critiques and Addresses'' (1873)
* ''American Addresses'' (1877)
* ''The Crayfish'' (1880)
* ''Science and Culture, and Other Essays'' (1881)
* ''Essays upon Some Controverted Questions'' (1892)
* ''Science and Hebrew Tradition'' (1893)
* ''Evolution and Ethics and Other Essays'' (1893)

==Further reading==
'''Biographies'''
* Ashforth, Albert. ''Thomas Henry Huxley''. Twayne, New York 1969.
* Ayres, Clarence. ''Huxley''. Norton, New York 1932.
* Bibby, Cyril. ''T.H. Huxley: Scientist, Humanist and Educator''. Watts, London 1959.
* Bibby, Cyril. ''Scientist Extraordinary: The Life and Work of Thomas Henry Huxley 1825–1895''. Pergamon, Oxford 1972.
* Clodd, Edward. ''Thomas Henry Huxley''. Blackwood, Edinburgh 1902.
* Desmond, Adrian. ''Huxley: vol 1 and 2''. Penguin, London 1998.
* Huxley, Leonard. ''Thomas Henry Huxley: A Character Sketch''. Watts, London 1920.
* Irvine, William. ''Thomas Henry Huxley''. Longmans, London 1960.
* Jensen, J. Vernon. ''Thomas Henry Huxley: Communicating for Science''. University of Delaware, Newark 1991. 
* Lyons, Sherrie L. ''Thomas Henry Huxley: The Evolution of a Scientist''. New York 1999.
* Mitchell, P. Chalmers. ''Thomas Henry Huxley: A Sketch of his Life and Work'' London 1901. Available at [https://www.gutenberg.org/ebooks/16935 Project Gutenberg].
* Voorhees, Irving Wilson. ''The Teachings of Thomas Henry Huxley''. Broadway, New York 1907.
* Huxley, Thomas Henry. ''Autobiography and Selected Essays''.  The Riverside Press Houghton Mifflin Company, Cambridge 1909.

==External links==
{{Commons category}}
{{Wikiquote}}
{{wikisource|works=or}}
* [http://embryo.asu.edu/pages/thomas-henry-huxley-1825-1895 Thomas H. Huxley] on the Embryo Project Encyclopedia
* [http://aleph0.clarku.edu/huxley/toc.html The Huxley File at Clark University]—Lists his publications, contains much of his writing.
* {{cite book|last=Stephen|first=Leslie|author-link=Leslie Stephen|title=Studies of a Biographer|volume=3|year=1898|publisher=Duckworth & Co.|location=London|pages=188–219|chapter=[[s:Studies of a Biographer/Thomas Henry Huxley|Thomas Henry Huxley]]}}
* [http://nla.gov.au/nla.party-869246 Huxley, Thomas Henry (1825–1895)] National Library of Australia, ''Trove, People and Organisation'' record for Thomas Huxley
* {{Gutenberg author |id=595| name=Thomas Henry Huxley}}
* [https://makingscience.royalsociety.org/s/rs/people/fst00041472 Science in the Making] Huxley's papers in the Royal Society's archives
* {{Internet Archive author |sname=Thomas Henry Huxley}}
* {{Librivox author |id=2741}}
* Huxley review: [http://darwin-online.org.uk/content/frameset?itemID=A166&viewtype=text&pageseq=1 Darwin on the origin of species] ''[[The Times]]'', 26 December 1859, p.&nbsp;8–9.
* Huxley review: [http://darwin-online.org.uk/content/frameset?itemID=A43&viewtype=image&pageseq=1 Time and life: Mr Darwin's "Origin of species."] ''Macmillan's Magazine'' 1: 1859 p.&nbsp;142–148.
* Huxley review: [http://darwin-online.org.uk/content/frameset?itemID=A32&viewtype=text&pageseq=1 Darwin on the origin of Species], ''[[Westminster Review]]'', 17 (n.s.) April 1860 p.&nbsp;541–570.

{{S-start}}
{{s-aca}}
{{s-bef|before = [[Thomas Wharton Jones]]}} 
{{s-ttl|title  = [[Fullerian Professor of Physiology]]|years  = 1855–1858}} 
{{s-aft|after  = [[Richard Owen]]}}
{{s-break}}
{{s-bef|before = [[John Marshall (surgeon)|John Marshall]]}} 
{{s-ttl|title  = [[Fullerian Professor of Physiology]]|years  = 1865–1869}} 
{{s-aft|after  = [[Michael Foster (physiologist)|Michael Foster]]}}
{{s-break}}
{{s-npo|pro}}
{{s-bef|before=[[William Spottiswoode]]}}
{{s-ttl|order=34th|title=[[Royal Society|President of the Royal Society]]|years=1883–1885}}
{{s-aft|after=[[Sir George Stokes, 1st Baronet|George Stokes]]}}
{{s-break}}
{{s-ach}}
{{s-bef|before= [[George Newport]]}}
{{s-ttl|title= [[Royal Medal]]|years=1852}}
{{s-aft|after= [[Charles Darwin]]}}
{{s-break}}
{{s-bef|before= [[Laurent-Guillaume de Koninck|L-G de Koninck]]}}
{{s-ttl|title= [[Wollaston Medal]]|years=1876}}
{{s-aft|after= [[Robert Mallet]]}}
{{s-break}}
{{s-bef|before= [[George Bentham]]}}
{{s-ttl|title= [[Clarke Medal]]|years=1880}}
{{s-aft|after= [[Frederick McCoy]]}}
{{s-break}}
{{s-bef|before= [[Alphonse de Candolle]]}}
{{s-ttl|title= [[Linnaean Medal]]|years=1890}}
{{s-aft|after= [[Jean-Baptiste Édouard Bornet|Jean-Baptiste Bornet]]}}
{{s-break}}
{{s-bef|before= [[Joseph Dalton Hooker]]}}
{{s-ttl|title= [[Darwin Medal]]|years=1894}}
{{s-aft|after= [[Giovanni Grassi]]}}
{{s-end}}

{{Copley Medallists 1851-1900}}
{{Royal Society presidents 1800s}}
{{GLS Presidents}}
{{Historical definitions of race}}
{{Rectors of the University of Aberdeen}}
{{Authority control}}

{{DEFAULTSORT:Huxley, Thomas Henry}}
[[Category:1825 births]]
[[Category:1895 deaths]]
[[Category:19th-century British anthropologists]]
[[Category:19th-century British zoologists]]
[[Category:19th-century English non-fiction writers]]
[[Category:19th-century English translators]]
[[Category:Academics of Imperial College London]]
[[Category:Alumni of Charing Cross Medical School]]
[[Category:Alumni of University College London]]
[[Category:Biological evolution]]
[[Category:British anthropologists]]
[[Category:British carcinologists]]
[[Category:British evolutionary biologists]]
[[Category:Burials at East Finchley Cemetery]]
[[Category:Charles Darwin]]
[[Category:Comparative anatomy]]
[[Category:Corresponding members of the Saint Petersburg Academy of Sciences]]
[[Category:Deans of the Royal College of Science]]
[[Category:English agnostics]]
[[Category:English anatomists]]
[[Category:English anthropologists]]
[[Category:English palaeontologists]]
[[Category:English sceptics]]
[[Category:English taxonomists]]
[[Category:English zoologists]]
[[Category:Fellows of the Linnean Society of London]]
[[Category:Fellows of the Royal Society]]
[[Category:Foreign associates of the National Academy of Sciences]]
[[Category:Fullerian Professors of Physiology]]
[[Category:Huxley family|Thomas Henry]]
[[Category:Knights of the Order of the Polar Star]]
[[Category:Members of the London School Board]]
[[Category:Members of the Privy Council of the United Kingdom]]
[[Category:Members of the Royal Netherlands Academy of Arts and Sciences]]
[[Category:Paleozoologists]]
[[Category:People from Ealing]]
[[Category:People involved in race and intelligence controversies]]
[[Category:Presidents of the Geological Society of London]]
[[Category:Presidents of the Royal Society]]
[[Category:Recipients of the Copley Medal]]
[[Category:Rectors of the University of Aberdeen]]
[[Category:Royal Medal winners]]
[[Category:Translators from German]]
[[Category:Victorian writers]]
[[Category:Wollaston Medal winners]]
[[Category:International members of the American Philosophical Society]]