Cordylus

Template:Short description Template:Automatic taxobox The genus Cordylus (Sauria: Cordylidae) includes a wide variety of species of small to medium spiny lizards from Africa, collectively called girdle-tailed lizards or girdled lizards. All are diurnal and ovoviviparous (live-bearing, without shelled eggs). Most species are rupicolous (rock-dwelling), while a few species are arboreal or live in burrows. They defend themselves with osteoderms (flat bony plates in the skin) and by quickly retreating into rock crevices or burrows. Many species live in groups, and males defend territories.

Cordylids are generally listed under CITES Appendix II. They are not necessarily threatened with extinction, but trade is controlled to prevent overexploitation. Some species of Cordylus have limited ranges and may be threatened with habitat destruction or over collecting for the pet trade.

Classification

Broadley (2006) recognized 47 species in the genus Cordylus, including eight species originally placed in the genus Pseudocordylus (P. fasciatus, P. langi, P. melanotus, P. microlepidotus, P. nebulosus, P. spinosus, P. subviridis, and P. transvaalensis) and Hemicordylus capensis. The greatest diversity of cordylids is in South Africa, with a few species found in Angola and eastern Africa as far north as Ethiopia. Other members of the Cordylidae are the genera Chamaesaura (a group of legless lizards from southern and eastern Africa) and Platysaurus. The sister group of Cordylidae is the plated lizards, family Gerrhosauridae.

In 2011, a study based on the molecular phylogeny of the family suggested several species traditionally included in Cordylus should be moved to other genera: Hemicordylus (for capensis and nebulosus), Karusasaurus (for jordani and polyzonus), Namazonurus (for campbelli, lawrenci, namaquensis, peersi and pustulatus), Ninurta (for coeruleopunctatus), Ouroborus (for cataphractus) and Smaug (for breyeri, giganteus, mossambicus, regius, vandami and warreni).<ref name= "Stanley et al 2011">Template:Cite journal</ref> This classification is supported by the Reptile Database where 21 species remain in Cordylus.<ref>Uetz, Peter et al. (2014). Template:EMBL genus www.reptile-database.org.</ref>

Genus Cordylus (sensu stricto):

• Angolan girdled lizard, Cordylus angolensis Template:Small
• Maasai girdled lizard, Cordylus beraduccii Template:Small
• Cape girdled lizard, Cordylus cordylus Template:Small
• Rooiberg girdled lizard, Cordylus imkeae Template:Small
• Limpopo girdled lizard, Cordylus jonesii Template:Small
• Machadoe's girdled lizard, Cordylus machadoi Template:Small
• Coastal spiny-tailed lizard, Cordylus macropholis Template:Small
• Marungu girdled lizard, Cordylus marunguensis Template:Small
• McLachlan's girdled lizard, Cordylus mclachlani Template:Small
• Mecula girdled lizard, Cordylus meculae Template:Small
• Dwarf girdled lizard, Cordylus minor Template:Small
• Kaokoveld girdled lizard, Cordylus namakuiyus Template:Small
• Black girdled lizard, Cordylus niger Template:Small

Cordylus niger, endemic to two small areas of South Africa.

• Nyika girdled lizard, Cordylus nyikae Template:Small
• Oelofsen's girdled lizard, Cordylus oelofseni Template:Small
• N’Dolondolo girdled lizard, Cordylus phonolithos Template:Small
• Rhodesian girdled lizard, Cordylus rhodesianus Template:Small
• Ethiopian girdled lizard, Cordylus rivae Template:Small
• East African spiny-tailed lizard, Cordylus tropidosternum Template:Small
• Ukinga girdled lizard, Cordylus ukingensis Template:Small
• Transvaal girdled lizard, Cordylus vittifer Template:Small

Nota bene: A binomial authority in parentheses indicates that the species was originally described in a genus other than Cordylus.

Formerly assigned in Cordylus (sensu lato):

• species in the genus Hemicordylus
• species in the genus Karusasaurus
• species in the genus Namazonurus
• Blue-spotted girdled lizard, Ninurta coeruleopunctatus
• Armadillo girdled lizard, Ouroborus cataphractus
• species in the genus Pseudocordylus
• species in the genus Smaug

Further reading

• Branch, Bill (2004). Field Guide to Snakes and other Reptiles of Southern Africa. Third Revised edition, Second impression. Sanibel Island, Florida: Ralph Curtis Books Publishing. 399 pp. Template:ISBN. (Genus Cordylus, pp. 185–186).
• Broadley DG (2006). Template:Cite web
• FitzSimons VF (1943). The Lizards of South Africa. Pretoria: Transvaal Museum Memoir.
• Laurenti JN (1768). Specimen medicum, exhibens synopsin reptilium emendatam cum experimentis circa venena et antidota reptilium austriacorum. Vienna: "Joan. Thom. Nob. de Trattnern". 214 pp. + Plates I–V. (Cordylus, new genus, p. 51). (in Latin).
• Reissig, Jens (2014). Girdled Lizards and their relatives: Natural History, Captive Care and Breeding. (Foreword by Aaron M. Bauer). Frankfurt am Main: Chimaira. 249 pp. Template:ISBN.
• Spawls S, Howell K, Drewes R, Ashe J (2002). A Field Guide to the Reptiles of East Africa. San Diego: Academic Press. 543 pp.

External links

• Tallbo
• Live Birth
• Sungrazers
• Armadillo Lizards