Hadar, Ethiopia

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Template:Short description Template:Use dmy dates Template:Distinguish Template:Infobox ancient site

Hadar or Hadar Formation (also spelled Qad daqar, Qadaqar; Afar "white [qidi] stream [daqar]")<ref name="Kalb-83">Jon Kalb Adventures in the Bone Trade (New York: Copernicus Books, 2001), p. 83</ref> is a paleontological fossil site located in Mille district, Administrative Zone 1 of the Afar Region, Ethiopia, 15 km upstream (west) of the A1 road's bridge across the Awash River (Adayitu kebele).<ref name="map">Template:Cite journal</ref>

It is situated on the southern edge of the Afar Triangle (part of East Africa's Great Rift Valley), along the left banks of the Awash River, between two minor tributaries, the eponymous Kada Hadar and the Kada Gona.<ref name="map" /> In 1972, Taieb organized a small exploratory reconnaissance of the Afar region to investigate more paleontological finds there. After six weeks of exploration, the party focused on the Hadar site.<ref name="DJohanson">Template:Cite journal</ref>

The site has yielded some of the most well-known hominin fossils, including "Lucy". These hominin fossils range in age from approximately 3.42 to 2.90 million years ago. These finds give us a greater understanding of hominin evolution during this period.

It is postulated that the specimens in the region were deposited by way of a large river system with associated crevasse channels/splays, deltas, and distributary channels, as well as periodic transgressions of paleolake Hadar located east of the research area (Aronson and Taieb, 1981, Tiercelin, 1986, Campisano and Feibel, in press) possibly related to geological activity or climatic cycles in at least the Kada Hadar Member (Yemane et al., 1996, Yemane, 1997, Campisano and Feibel, in press)."

According to Jon Kalb, early maps show caravan routes passing within 10 to 15 km of Hadar but not through it. The British explorer L.M. Nesbitt passed 15 km west of Hadar in 1928.<ref name="Kalb-83" /> Template:TOC limit

Geology

File:Ethiopia Afar Zone1.PNG
Administrative Zone 1 (Afar Region), Ethiopia

The region's rocks consist mainly of mudstones, siltstones, fine-grained sandstones and volcanic tuffs. The region of Hadar has been divided into four geologic members — Basal (~3.8–3.42 Ma), Sidi Hakoma (~3.42–3.26 Ma), Denen Dora (~3.26–3.2 Ma), and Kada Hadar (<~3.2 Ma)—with three tuffs (Sidi Hakoma Tuff [SHT], Triple Tuff [TT] and Kada Hadar Tuff [KHT]) separating the four members.

The Sidi Hakoma member tends towards high rainfall and low seasonality. The overlying Denan Dora Member was a grassland habitat. Finally, the Kada Hadar Member was an even more open and arid habitat, as seen in the high abundance of antilopines, which frequent these types of terrains.<ref name="DJohanson" />

Paleontology

The first paleo-geological explorations of the Hadar area were conducted by Maurice Taieb. He found Hadar in December 1970 by following the Ledi River, which originates in the highlands north of Bati to empty into the Awash River. Taieb recovered several fossils in the area and led a party back to Hadar in May 1972. In October 1973, 16 individuals with the International Afar Research Expedition (IARE) arrived at Hadar and camped there for two months during which the first hominin fossil was found. (Taieb claims in his 1985 book Sur la Terre des premiers Hommes to have discovered the Hadar fields in 1968, but Kalb argues that claim to be incorrect.)<ref>Template:Cite bookTemplate:Pn</ref> The IARE party examined a series of sedimentary layers called the Hadar Formation, which was dated to the late Pliocene to early Pleistocene epochs (3.5 to 2.3 million years ago).<ref name="feibel">Template:Cite conference</ref>

Discovery of Lucy

File:Reconstruction of the fossil skeleton of "Lucy" the Australopithecus afarensis.jpg
Lucy", a 3.2-million-year-old Australopithecus afarensis fossil discovered in Hadar

The anthropologist Donald Johanson, a member of the 1973 expedition to Hadar, returned the next year and discovered the fossil hominin "Lucy" in the late fall of 1974.<ref name=":0">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> He spotted a right proximal ulna in a gully, followed by an occipital bone, a femur, some ribs, a pelvis, and a lower jaw. Within two weeks, nearly 40% of the hominoid skeleton had been identified and cataloged.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Lucy is the most famous fossil to have been found at Hadar. Lucy is among the oldest hominin fossils ever discovered<ref name=":0" /> and was later given the taxonomic classification Australopithecus afarensis. (The name 'Lucy' was inspired by the song "Lucy in the Sky with Diamonds" by The Beatles, which happened to be playing on the radio at base camp.)

Specimens and inferences

In 1975, Donald Johanson made another discovery at a nearby site in Hadar: 216 specimens from approximately 17 individuals, most likely related and varying in age, called AL 333 (colloquially referred to as the "First Family").

About thirty years later in nearby Dikika, another Australopithecus afarensis fossil skeleton was found in a separate outcrop of the Hadar Formation across the Awash River from Hadar. The skeleton is of a three-year-old girl later named "Selam," which means peace in Amharic Ethiopian languages.

In 1973 and 1974 when the first anatomical discoveries were made, their size and shape pointed towards a variety of taxa, but further research has confirmed that only one hominin taxon is present here. The first find there was a fossil knee joint estimated to date from 3.4 million years ago. Since then, the Hadar research area has yielded 370 specimens of A. afarensis, one specimen of Homo, and 7571 additional vertebrate specimens.

The specimens recovered display a variety of different primitive cranial post features, which indicate A. afarensis is distinct from other species of Australopithecus: small cranial capacity, palate similar to African apes (parallel tooth rows, shallow, long from front to back, narrow from side to side), primitive occipital, basal cranium anatomy, high frequency of unicuspid third premolars, prognathic face, and primitive mandibular anatomy. Postcranially, the pelvis, knee, ankle, and foot indicate habitual, terrestrial bipedalism, but ape-like curved finger and foot bones are retained ancestral ape-like features.<ref name="DJohanson" />

Paleofauna

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Although Australopithecus is well-known for its discovery since the 1970s, other specimens from the Hadar Formation contain several fossil remains of artiodactyls, perissodactyls, carnivorans, proboscideans, and other African species that are well preserved. The bovids found in the formation included the Aepycerotini (Aepyceros), Alcelaphini (Damalborea and Parmularius), Antilopini (Gazella), Bovini (Ugandax and Pelorovis[?]), Caprini (Budorcas), Cephalophini, Hippotragini (Oryx), Neotragini (Raphicerus[?] and Madoqua), Reduncini (Kobus), and Tragelaphini (Tragelaphus).<ref name="Tragelaphini">Template:Cite journal</ref><ref name="Bovidae">Template:Cite journal</ref><ref>Template:Cite journal</ref> Artiodactyls outside the bovid family were present within the formation as well, namely the giraffids (Giraffa and Sivatherium), Hippopotamidae (Hippopotamus), and suids (Kolpochoerus, Notochoerus, and Nyanzachoerus).<ref name = suidae>Template:Cite journal</ref> While a definitive list of carnivorans found within the Hadar Formation has yet to be compiled, confirmed genera that were found within the Hadar Formation include canids (Canis and Nyctereutes), felids (Dinofelis,<ref name = dinofelis>Template:Cite journal</ref> Leptailurus, Felis, Homotherium, and Panthera), hyaenids (Chasmaporthetes, Ikelohyaena, Crocuta, Hyaena, and cf. Pliocrocuta),<ref name = hyaena>Template:Cite journal</ref> herpestids (Herpestes and cf. Helogale), mustelids (Mellivora, Enhydriodon,<ref name = enhydriodon>Template:Cite journal</ref> and cf. Poecilogale), and the viverrid (cf. Civettictis). Mammals within the formation outside the artiodactyl and carnivoran families include a bat (indeterminate), the leporid (Lepus), the equid (Eurygnathohippus),<ref name = equid>Template:Cite book</ref> rhinoceroses (Ceratotherium and Diceros),<ref name = rhinoceros>Template:Cite journal</ref> old world primates (Parapapio, Theropithecus, and Cercopithecoides), proboscideans (the deinothere Deinotherium and elephants Elephas, Loxodonta, and Mammuthus) <ref name = elephantoids>Template:Cite journal</ref><ref name = proboscidean>Template:Cite journal</ref><ref>Template:Cite journal</ref> old world porcupines (Hystrix and Xenohystrix),<ref name = hystrix>Template:Cite journal</ref> murid rodents (Gerbilliscus, Acomys, Golunda, Oenomys, Praomys, Saidomys, Millardia, and Mus),<ref name = Murid>Template:Cite journal</ref> the spalacid Tachyoryctes, a squirrel indet., and an aardvark species. Taxons within other classes are present within the Hadar Formation as well, such as birds (Plectropterus, Balearica, Anhinga, and Struthio) and reptiles (Crocodylus, Python, Varanus, and Bitis).<ref>Template:Cite book</ref><ref name = carnivora>Template:Cite journal</ref><ref name="Giraffa">Template:Cite journal</ref><ref name = Cercopithecidae>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref name="Reed">Template:Cite journal</ref><ref>Template:Cite journal</ref> Template:Paleobiota-key-compact

Mammals

Artiodactyls

Bovidae
Bovids of the Hadar Formation
Taxa Species Presence Material Notes Images
Aepyceros A. datoadeni Kada Hadar Member at Hadar Near-complete skull lacking parts of horn cores (AL 787-1 KH1)<ref name = Bovidae/> An impala
Beatragus B. sp. Detached braincase with two horn cores<ref name = Bovidae/> A hirola
Budorcas B. churcheri<ref>Template:Cite book</ref> Sida Hakoma Member.<ref name = Bovidae/> Complete skull with horns (AL 136-5 DD2)<ref name = Bovidae/> An extinct takin
Damalops D. sidihakomai<ref>Template:Cite journal</ref> Sidi Hakoma Member AL 208–7, a skull with horn cores, right P4–M3, and left M2–3. An extinct genus of Alcelaphinae
Damalborea D. elisabethae Basal and Sidi Hakoma members<ref name = Bovidae/> A skull with horn cores (AL 208-07 SH3)<ref name = Bovidae/> An extinct genus of Alcelaphinae
D. grayi Denen Dora and Kada Hadar members<ref name = Bovidae/> Multiple postcranial fragments, such as horn cores, maxillae, and mandibles<ref name = Bovidae/>
Gazella G. harmonae Kada Hadar Member at Hadar<ref name = Bovidae/> Skull remains, such as isolated teeth, maxillae, frontlets, and occipital bones (AL 444-16)<ref name = Bovidae/> Extinct relatives of modern gazelles
G. cf. janenschi Denen Dora Member Horn cores (AL 302-6 DD)<ref name = Bovidae/>
G. cf. praethomsoni Denen Dora and Kada Hadar members Few Horn core specimens<ref name = Bovidae/>
Hippotragini Praedamalis deturi Kada Hadar, Sida Hakoma, and Denen Dora members at Hadar Multiple specimens based on horn cores and lower teeth<ref name = Bovidae/> Ancestor of modern oryx
Kobus K. hadarensis Sidi Hakoma, Kada Hadar and Denen Dora member Extinct relatives of antelopes
K. oricornus Denen Dora and Kada Hadar members Several specimens of braincase and horn core<ref name = Bovidae/>
Madoqua M. sp. Relative of dik-dik
Raphicerus R. sp. Kada Hadar and Sidi Hakoma members Mandibles and horn cores<ref name = Bovidae/> Relative of Neotragini
Parmularius P. pachyceras Sidi Hakoma and Kada Hadar members<ref name = Bovidae/> Braincases and horn cores<ref name = Bovidae/> Large extinct alcelphines related to topis and hartebeests
Pelorovis P. sp.
Tragelaphus T. lockwoodi Basal and Sidi Hakoma members Braincase with complete horn cores (AL 142-3 SH2)<ref name = Bovidae/> Extinct relatives of spiral-horned antelopes
T. rastafari Basal, Sidi Hakoma, Denen Dora, and Kada Hadar members Several horn core specimens
Ugandax U. coryndonae Denen Dora and Sidi Hakoma members at Hadar Complete partial skull and horns (AL 194-1 DD1)<ref name = Bovidae/> Ancestor of Cape buffalo
File:Ugandax gautieri 2.jpg
Giraffidae
Giraffids of the Hadar Formation
Taxa Species Presence Material Notes Images
Giraffa G. jumae Sidi Hakoma and Denen Dora members Partial teeth, vertebrae, ossicones, limb bones, and mandibles<ref name = Giraffa/> Extinct relatives of modern giraffe
G. stillei Sidi Hakoma and Denen Dora members Partial teeth, ossicones, limb bones, and mandibles<ref name = Giraffa/>
Sivatherium S. maurusium Denen Dora, Kada Hadar, and Sidi Hakoma members Partial mandibles and horns<ref name = Giraffa/> Okapi-like giraffid
File:MEPAN Sivatherium.jpg
Hippopotamidae
Hippopotamids of the Hadar Formation
Taxa Species Presence Material Notes Images
Hexaprotodon H. protoamphibius Denen Dora Member Partial skull and teeth Extinct relatives of modern pygmy hippo
H. coryndoni Sidi Hakoma Member<ref name = Reed/> Partial skull and teeth
Hippopotamus H. afarensis Basal, Sidi Hakoma, Kada Hadar, and Denen Dora members<ref name = Reed/> Complete postcranial skull and mandibles Extinct subspecies related to common hippopotamus, formally named Trilobophorus afarensis
Suidae
Suids of the Hadar Formation
Taxa Species Presence Material Notes Images
Kolpochoerus K. afarensis Kada Hadar, Sidi Hakoma, and Denen Dora members<ref name = suidae/> Postcranial skulls, teeth fragments, and mandibles Extinct relative of the pig family
File:Kolpochoerus limnetes 3.jpg
Notochoerus N. eulius Kada Hadar, Lower and Upper Sidi Hakoma, and Denen Dora members<ref name = suidae/> Postcranial skull, teeth, and mandibles A Tetraconodontinae pig.
Nyanzachoerus N. kanamensis Basal and Sidi Hakoma members<ref name = Reed/> Complete cranium, mandibles, and dentition A Tetraconodontinae pig.
N. pattersoni Basal, Lower and Upper Sidi Hakoma, and Denen Dora members<ref name = suidae/> Complete skulls, mandibles, and dentition<ref name = suidae/>

Carnivorans

Canidae
Canids of the Hadar Formation
Taxa Species Presence Material Notes Images
Nyctereutes N. lockwoodi Sidi Hakoma and Denen Dora members at Dikika<ref name = carnivora/> Nearly complete skull (DIK-31-1)<ref>Template:Cite journal</ref> A raccoon dog
Felidae
Felids of the Hadar Formation
Taxa Species Presence Material Notes Images
Dinofelis D. aronoki Kada Hadar Member<ref name = dinofelis/> Postcranial member: distal radius (AL-363-20)<ref name = dinofelis/> A machairodontine felid
File:Dinofelis15DB.jpg
D. petteri Sidi Hakoma and Denen Dora members<ref name = dinofelis/> Postcranial member: partial humerus (AL 168-30), proximal ulna (AL 262-9), and right astragalus (Al 173-11)<ref name = dinofelis/>
Felis F. sp. Kada Hadar, Sidi Hakoma, and Denen Dora members A small cat relative
Homotherium H. hadarensis Denen Dora Member<ref name = carnivora/> Complete cranium skull, mandible fragments (DIK-96-1), teeth, and forelimbs<ref name = carnivora/> A machairodontine felid
Megantereon M. sp. Kada Hadar, Sidi Hakoma, and Denen Dora members A machairodontine felid
Leptailurus L. sp. A serval cat
Panthera P. sp. Sidi Hakoma Member Relatives of lions
Herpestidae
Herpestids of the Hadar Formation
Taxa Species Presence Material Notes Images
Herpestes H. sp. Dikika site Tooth fragments A mongoose
Hyaenidae
Hyaenids of the Hadar Formation
Taxa Species Presence Material Notes Images
Chasmaporthetes C. sp. Sidi Hakoma and Denen Dora members at Hadar<ref name = hyaena/> A hunting hyena
Crocuta C. dietrichi Sidi Dakoma , Denen Dora, and Basal members at Hadar<ref name = hyaena/> Fragmented mandibles (DIK-32-1)<ref name = carnivora/><ref name = hyaena/> Extinct relatives of modern hyena
C. eturono Kada Hadar Member Partial maxilla (DIK-73-1)<ref name = carnivora/>
Hyaena H. sp.
Ikelohyaena I. abronia Sidi Hakoma Member at Hadar<ref name = hyaena/> Early member of the Parahyaena lineage and the earliest close relative of the modern striped hyena
Percrocuta P. sp.
Pliocrocuta P. cf. perierri Sidi Hakoma and Denen Dora members An early hyaena relative, synonym named Pachycrocuta cf. perrieri
Mustelidae
Mustelids of the Hadar Formation
Taxa Species Presence Material Notes Images
Enhydriodon E. dikikae Basal and Sidi Hakoma members at Dikika site<ref name = enhydriodon/><ref name = carnivora/> Partial skull (DIK-56-9) and humerus (DIK-78-1)<ref name = enhydriodon/><ref name = carnivora/> A giant river otter
Lutra L. hearsti Lower part of Sidi Hakoma Member at Dikika Upper teeth fragments (DIK-50–35)<ref name = carnivora/> Extinct relative of otters
Mellivora M. sp. A honey badger
Poecilogale P. sp. Sidi Hakoma Member
Viverridae
Viverrids of the Hadar Formation
Taxa Species Presence Material Notes Images
Civettictis C. sp. A civet

Perissodactyls

Equidae
Equids of the Hadar Formation
Taxa Species Presence Material Notes Images
Eurygnathohippus E. afarense Kada Hadar Member Partial skull (AL 363-18) and mandibles<ref name = equid/> A hipparionine horse.
E. hasumense Denen Dora Member Partial skull (AL 340-8) and mandibles<ref name = equid/>
Rhinocerotidae
Rhinocerotids of the Hadar Formation
Taxa Species Presence Material Notes Images
Ceratotherium C. mauritanicum Sidi Hakoma Member at Lower Awash of Dikika Complete cranium skull (AL-129-25)<ref name = rhinoceros/> Ancestor of modern white rhinoceros
Diceros D. praecox Sidi Hakoma Member at Lower Awash of Dikika Incomplete cranium skull fragments and jawbones<ref name = rhinoceros/> Direct ancestor of modern black rhinoceros, formally named Ceratotherium praecox

Primates

Cercopithecidae
Cercopithecids of the Hadar Formation
Taxa Species Presence Material Notes Images
Cercopithecoides C. meaveae Sidi Hakoma Member Partial skeleton (A.L. 2-64 and A.L. 222-14)<ref name = Cercopithecidae/> A colobine monkey
Rhinocolobus R. turkanaensis Sidi Hakoma and Denen Dora members Preserved mandibles and teeth fragments<ref name = Cercopithecidae/> Extinct relative of colobus monkey
Parapapio P. cf. jonesi Sidi Hakoma and Kada Hadar members Complete postcranial skull, partial limbs, teeth fragments, and mandibles<ref name = Cercopithecidae/> A prehistoric baboon
Theropithecus oswaldi<ref>Template:Cite journal</ref> T. o. darti Basal, Denen Dora, Kada Hadar, and Sidi Hakoma members Postcranial skulls differentiated between male and female<ref name = Cercopithecidae/> Extinct relatives of gelada baboons
T. o. ecki Denen Dora Member Female cranium skull and mandible
T. o. oswaldi Sidi Hakoma and Kada Hadar members Postcranial skulls differentiated between male and female<ref name = Cercopithecidae/>
Hominidae
Hominids of the Hadar Formation
Taxa Species Presence Material Notes Images
Australopithecus A. afarensis Basal, Denen Dora, Kada Hadar, and Sidi Hakoma members Partial skeletons featuring small cranial capacity, primitive occipitals, basal cranium anatomy, high frequency of unicuspid third premolars, prognathic faces, and primitive mandibular anatomy An australopithecine hominid
File:Reconstruction of the fossil skeleton of "Lucy" the Australopithecus afarensis.jpg

AL 444-2 skull
AL 444-2 skull

Proboscideans

Deinotheriidae
Deinotheriids of the Hadar Formation
Taxa Species Presence Material Notes Images
Deinotherium D. bozasi Denen Dora, Kada Hadar, and Sidi Hakoma members Complete cranium skull and teeth fragments A large extinct elephant-like proboscidean
File:Deinotherium bozasi JG.png
Elephantidae
Elephantids of the Hadar Formation
Taxa Species Presence Material Notes Images
Elephas E. ekorensis Sidi Hakoma and Denen Dora members at Hadar and Dikika<ref name = proboscidean/> Mandibles and molar fragments Extinct lineage related to Asian elephant
Loxodonta L. adaurora Denen Dora and Sidi Hakoma members at Hadar<ref name = elephantoids/> Mandibles and molar fragments Extinct subspecies of modern African elephant
L. exoptata Denen Dora Member Undescribed molars<ref name = elephantoids/>
Mammuthus M. sp. Sidi Hakoma and Denen Dora Member. Undescribed molars<ref name = elephantoids/> A mammoth relative
Palaeoloxodon recki P. r. brumpti Sidi Hakoma Member<ref name = proboscidean/> Mandibles and molar fragments Extinct relative of African elephant
File:Elaphus recki 2.png

Rodents

Rodents of the Hadar Formation
Taxa Species Presence Material Notes Images
Acomys A. cf. lavocati Mandible specimens<ref name = Murid/> Relatives of Spiny mice.
A. cf. lemniscomys Right mandible and broken tooth root<ref name = Murid/>
Gerbilliscus G. sp. Right maxillary fragment with M2-3 and right mandible fragment with M1-3<ref name = Murid/> Relative of gerbil species
Golunda G. gurai Murine rodent closely related to the Indian bush rat
Hystrix H. cf. makapanensis Sidi Hakoma Member<ref name = hystrix/> An Old World porcupine
Mastomys M. cf. minor Partial skull and mandibles<ref name = Murid/> Extinct relative of the multimammate mouse
Millardia M. coppensi Extinct relatives of soft-furred rat
M. taiebi
Mus M. indet. Unknown relative of Murini
Oenomys O. tiercelini Extinct relative of modern rusty-nosed rat
Pelomys P. cf dietrichi Mandibles A groove-toothed swamp rat
Praomys P. sp. Sidi Hakoma Member A.L. 894-35993, a right mandible with M1-3<ref name= Murid/> A Muridae species
Saidomys S. afarensis Sidi Hakoma Member Extinct relative of a muroid mouse
Tatera T. sp. Sidi Hakoma and Denen Dora members Extinct relative of the Indian gerbil
Thallomys T. cf. quadrilobatus Complete skull with both mandibles<ref name = Murid/> Extinct relative of the rat family endemic to Africa
Thryonomys T. swinderianus A cane rat
Xenohystrix X. cf. crassidens Sidi Hakoma and Denen Dora members An old-world porcupine
Xerus X. sp. Kada Hadar Member An unstriped ground squirrel

Birds

Birds of the Hadar Formation
Taxa Species Presence Material Notes Images
Anhinga A. hadarensis<ref>Template:Cite journal</ref> An extinct darter
Balearica B. sp.
Plectropterus P. sp.
Struthio S. sp.

Reptiles

Reptiles of the Hadar Formation
Taxa Species Presence Material Notes Images
Bitis B. sp.
Crocodylus C. sp.
Python P. sp.
Varanus V. sp.

See also

References

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