Holometabola
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Holometabola (from Ancient Greek Template:Wikt-lang "complete" + Template:Wikt-lang "change"), also known as Endopterygota (from Template:Wikt-lang "inner" + Template:Wikt-lang "wing" + Neo-Latin Template:Wikt-lang "-having"), is a supra-ordinal clade of insects within the infraclass Neoptera that go through distinctive larval, pupal, and adult stages. They undergo a radical metamorphosis, with the larval and adult stages differing considerably in their structure and behaviour. This is called holometabolism, or complete metamorphism.
Evolution
The Holometabola constitute the most diverse insect superorder, with over 1 million living species divided between 11 orders, containing insects such as butterflies, flies, fleas, bees, ants, and beetles.<ref name="Beutel">Template:Cite journal</ref>
The earliest holometabolan fossils date from the Carboniferous.<ref>Template:Cite journal</ref>
The Holometabola are sometimes divided into three assemblages: Neuropterida (Neuroptera, Megaloptera, Raphidioptera, Strepsiptera and Coleoptera), Hymenopteroida (Hymenoptera), and Panorpida (Siphonaptera, Diptera, Trichoptera, Lepidoptera and Mecoptera).
Molecular analysis has clarified the group's phylogeny, as shown in the cladogram.<ref name=Kjer>Template:Cite journal</ref>
Description
The Endopterygota are distinguished from the Exopterygota ("external winged forms") by the way in which their wings develop. Endopterygota ("internal winged forms") develop wings inside the body and undergo an elaborate metamorphosis, holometabolism, involving a pupal stage. The Exopterygota develop wings on the outside of their bodies and do not go through a pupal stage. The Exopterygota are not a natural group (they are paraphyletic).<ref name="RES">Template:Cite web</ref>