Sceptridium

Sceptridium is a genus of seedless vascular plants in the family Ophioglossaceae,<ref name="Christenhusz_2011b">Template:Cite journal</ref> closely allied to (and often included as a subgenus<ref>subgenus Tropicos.org. Missouri Botanical Garden. 16 Jan 2012</ref> of) the genus Botrychium (the moonworts and grapeferns). It is also closely related to the genus Botrypus (the rattlesnake fern, often treated as the subgenus Osmundopteris under Botrychium). Sceptridium species are commonly called the grape-ferns.

These plants are small with fleshy roots, and reproduce by spores shed into the air. They differ from the moonworts in having at least some sterile fronds (all fronds in Botrychium are spore-bearing), and in the fronds being bi- or tri-pinnate (Botrychium are single pinnate, or rarely bipinnate); and from Botrypus in being evergreen, or at least winter-green (Botrypus are deciduous) and having the non-spore-bearing part of the frond long-stalked (short-stalked in Botrypus).

Taxonomy

The Smith et al. classification of 2006, based on molecular phylogeny, lumped Sceptridium within Botrychium, in the Ophioglossaceae,<ref name=Smith-2006>Template:Cite journal</ref> which was continued in the Christenhusz et al. classification of 2011<ref name=Christenhusz-2011>Template:Cite journal</ref> and the Christenhusz and Chase classification of 2014.<ref name=ChriChas14>Template:Cite journal</ref> It is recognized as a distinct genus in Ophioglossaceae by the Pteridophyte Phylogeny Group classification of 2016 (PPG I).<ref name=PPGI>Template:Citation</ref>

Phylogeny

Phylogeny of Sceptridium<ref>Template:Cite journal</ref><ref>Template:Cite web</ref> Template:Clade

Unassigned species:

Botrychium rugulosum

• S. × argutum Sahashi
• S. australe (Brown) Lyon (parsley fern)<ref>Template:Cite journal</ref><ref>B. australe Plants for a Future 26 Dec 2011</ref>
• S. biforme (Col.) Lyon (fine-leaved parsley fern)<ref>B. biforme International Plant Names Index (IPNI) 02 Jan 2012</ref><ref>S. biforme International Plant Names Index (IPNI) 02 Jan 2012</ref>
• S. × elegans Sahashi
• S. javanicum Sahashi
• S. jenmanii (Underwood) Lyon (Alabama grape-fern)<ref>B. jenmanii Flora of North America, www.eFloras.org 26 Dec 2011</ref><ref>B. jenmanii Tropicos.org. Missouri Botanical Garden. 26 Dec 2011</ref><ref>B. alabamense Tropicos.org. Missouri Botanical Garden. 27 Dec 2011</ref>
• S. × longistipitatum Sahashi
• S. × pulchrum Sahashi
• S. rugulosum (Wagner) Skoda & Holub (ternate grape-fern, St. Lawrence grape fern, rugulose grape fern)<ref name=mich/><ref>B. rugulosum Tropicos.org. Missouri Botanical Garden. 26 Dec 2011</ref><ref name=botrug>B. rugulosum Robert W. Freckmann Herbarium, University of Wisconsin - Stevens Point 27 Dec 2011</ref>
• S. × silvicola (Sahashi) Ebihara
• S. subbifoliatum (Brack.) Lyon (island grape-fern)<ref>Template:PLANTS</ref><ref>B. subbifoliatum International Plant Names Index 27 Dec 2011</ref><ref>B. subbifoliatum ITIS taxonomy 26 Dec 2011</ref>
• S. underwoodianum (Maxon) Lyon<ref>B. underwoodianum Tropicos.org. Missouri Botanical Garden. 27 Dec 2011</ref><ref>S. underwoodianum Tropicos.org. Missouri Botanical Garden. 27 Dec 2011</ref>