Stachys
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Stachys is a genus of plants, one of the largest in the mint family Lamiaceae.<ref name="harley2004">Harley, R. M., et al. 2004. "Labiatae". pages 167–275. In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). The Families and Genera of Vascular Plants volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. Template:ISBN</ref> Estimates of the number of species vary from about 300<ref name="harley2004" /> to about 450.<ref name="mabberley2008">Mabberley, D. J. 2008. Mabberley's Plant-Book third edition (2008). Cambridge University Press: UK.</ref> Plants of the World Online accepts 373 species.<ref name = powo/> Stachys is in the subfamily Lamioideae<ref name="harley2004" /> and its type species is Stachys sylvatica.<ref name="ingstachys">Stachys In: Index Nominum Genericorum. In: Regnum Vegetabile (see External links below).</ref> The precise extent of the genus and its relationship to other genera in the subfamily are poorly known.
Range and naming
The distribution of the genus covers Europe, Asia, Africa, Australasia and North America. Common names include hedgenettle,<ref>Template:PLANTS</ref> heal-all, self-heal, woundwort, betony, Monty plant and lamb's ears. Wood betony, S. officinalis, was the most important medicinal herb to the Anglo-Saxons of early medieval England, and was used for many medicinal purposes from Ancient Roman times to the Early Modern period.
Stachys was named by Linnaeus in Species Plantarum in 1753.<ref name="linnaeus1753">Carolus Linnaeus. 1753. Species Plantarum 2:580. Laurentii Salvii. (see External Links below).</ref> The name is derived from the Greek word σταχυς (stachys), meaning "an ear of grain",<ref name="quattrocchi2000">Umberto Quattrocchi. 2000. CRC World Dictionary of Plant Names volume I, page 91. CRC Press: Boca Raton; New York; Washington, DC;, USA. London, UK. Template:ISBN (set). (see External links below).</ref> and refers to the fact that the inflorescence is often a spike. The name woundwort derives from the past use of certain species in herbal medicine for the treatment of wounds.
Human uses
The Chinese artichoke or Crosne (S. affinis), is grown for its edible tuber.<ref name="mabberley2008" /> Several species are cultivated as ornamentals. Woolly betony (S. byzantina) is a popular decorative garden plant. Wood betony (S. officinalis) was historically a highly valued medicinal plant.
Use by other species
Stachys species are used as food plants by the larvae of some Lepidoptera species, including the moths Coleophora auricella, C. lineolea, and C. wockeella, all recorded on S. officinalis. They are also widely used by the European wool carder bee (Anthidium manicatum), which scrape the hairs from the plant in order to use them for building their nests.<ref name=alpha>Template:Cite journal</ref>
Description
Stachys is a genus of shrubs and annual or perennial herbs. The stems vary from Template:Cvt tall, with simple, opposite, triangular leaves, Template:Cvt long with serrate margins. In most species, the leaves are softly hairy. The flowers are Template:Cvt long, clustered in the axils of the leaves on the upper part of the stem. The corolla is 5-lobed with the top lobe forming a 'hood', varying from white to pink, purple, red or pale yellow.
Circumscription
The distinction between Stachys and other genera is unclear and has varied from one author to another. In 2002, a molecular phylogenetic study showed that Stachys officinalis is not closely related to the rest of the genus.<ref name="lindqvist2002">Lindqvist, C. and V. A. Albert. 2002. Origin of the Hawaiian endemic mints within North American Stachys (Lamiaceae). American Journal of Botany 89(10), 1709–24.</ref> This study also found six other genera to be embedded within Stachys as it is currently circumscribed. The embedded genera are Prasium, Phlomidoschema, Sideritis, Haplostachys, Phyllostegia, and Stenogyne.
Fossil record
†Stachys pliocenica fossil seeds are known from Upper Miocene strata of Bulgaria and Pliocene strata of south-eastern Belarus. The fossil seeds are similar to the seeds of Stachys cretica.<ref>The Pliocene flora of Kholmech, south-eastern Belarus and its correlation with other Pliocene floras of Europe by Felix Yu. Velichkevich and Ewa Zastawniak - Acta Palaeobot. 43(2): 137–259, 2003</ref>
Diversity
Plants of the World Online accepts 373 species.<ref name = powo/> Selected species include:
- Stachys affinis Bunge – Chinese artichoke
- Stachys ajugoides Benth. – bugle hedgenettle
- Stachys alabamica B.R.Keener & L.J.Davenp. – Alabama hedgenettle
- Stachys albens A.Gray – white hedgenettle, whitestem hedgenettle
- Stachys albicaulis Lindl.
- Stachys alpina L. – alpine woundwort
- Stachys annua (L.) L. – annual woundwort
- Stachys arvensis L. – staggerweed, field woundwort
- Stachys bullata Benth.
- Stachys byzantina K.Koch – woolly betony, lamb's ear
- Stachys candida Bory & Chaub.
- Stachys chamissonis Benth. – great hedge nettle, coastal hedge nettle
- Stachys chamissonis var. cooleyae (A.Heller) G.A.Mulligan & D.B.Munro – Cooley's hedge nettle
- Stachys chrysantha Boiss. & Heldr.
- Stachys citrina Benth.
- Stachys clingmanii Small – Clingman's hedge nettle
- Stachys coccinea Ortega
- Stachys corsica Pers.
- Stachys cretica L.
- Stachys debilis Kunth
- Stachys ehrenbergii Boiss.
- Stachys floridana Benth. – Florida hedgenettle, Florida betony
- Stachys germanica L. – downy woundwort
- Stachys glutinosa L.
- Stachys hispida Pursh – rough hedge-nettle
- Stachys hyssopifolia Michx.
- Stachys iva Griseb.
- Stachys lavandulifolia Vahl
- Stachys libanotica Benth.
- Stachys manantlanensis B.L.Turner
- Stachys mexicana Benth. – Mexican hedge nettle
- Stachys milanii Magnier
- Stachys nelsonii B.R.Keener & L.J.Davenp. – Nelson's hedgenettle
- Stachys palustris L. – marsh woundwort, marsh hedge-nettle
- Stachys pumila Banks & Sol.
- Stachys pycnantha Benth.
- Stachys recta L. – yellow woundwort
- Stachys riederi Cham.
- Stachys rigida Benth. – rough hedgenettle
- Stachys sprucei Briq.
- Stachys stricta Greene
- Stachys sylvatica L. – hedge woundwort
- Stachys tenuifolia Willd. – smooth hedgenettle
Formerly placed here
- Agastache foeniculum (Pursh) Kuntze (as S. foeniculum Pursh)
- Leonurus japonicus Houtt. (as S. artemisia Lour.)
- Stachys monieri (Gouan) P.W.Ball, now in the synonymy of Betonica officinalis
- Betonica macrantha K.Koch (as S. macrantha (K.Koch) Stearn)
- Betonica officinalis L. (as S. officinalis (L.) Trevis.)