Tropism

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File:Bellis perennis - Gaenseblume4.jpg
Daisies (Bellis perennis) facing the Sun after opening in the morning showing heliotropism
File:Phycomyces3.JPG
Phycomyces, a fungus, exhibiting phototropism

In biology, a tropism is a phenomenon indicating the growth or turning movement of an organism, usually a plant, in response to an environmental stimulus.<ref>Template:Cite book</ref> In tropisms, this response is dependent on the direction of the stimulus (as opposed to nastic movements, which are non-directional responses). Tropisms are usually named for the stimulus involved; for example, a phototropism is a movement to the light source, and an anemotropism is the response and adaptation of plants to the wind.<ref>Template:Cite web</ref>

Tropisms occur in three sequential steps. First, there is a sensation to a stimulus. Next, signal transduction occurs. And finally, the directional growth response occurs.

Tropisms can be regarded by ethologists as taxis (directional response) or kinesis (non-directional response).

The Cholodny–Went model, proposed in 1927, is an early model describing tropism in emerging shoots of monocotyledons, including the tendencies for the stalk to grow towards light (phototropism) and the roots to grow downward (gravitropism). In both cases, the directional growth is considered to be due to asymmetrical distribution of auxin, a plant growth hormone.<ref name="Plant Cell" />

Template:AnchorThe term "tropism" (Template:Ety) is also used in unrelated contexts. Viruses and other pathogens affect what is called "host tropism", "tissue tropism", or "cell tropism"; in which case tropism refers to the way in which different viruses/pathogens have evolved to preferentially target specific host species, specific tissue, or specific cell types within those species. In English, the word tropism is also used to indicate an action done without cognitive thought: However, "tropism" in this sense has a proper, although non-scientific, meaning as an innate tendency, natural inclination, or propensity to act in a certain manner towards a certain stimulus.

Types

Tropisms can be distinguished according to the orientation with respect to the direction of the stimulus. They can commonly be either positive (towards the stimulus) or negative (away from it).<ref name=":0">Template:Cite web</ref> Both of these are orthotropic,<ref>Template:Cite OED</ref> and can be contrasted with tropisms that are diatropic (perpendicular to the stimulus) or plagiotropic (at an oblique angle).<ref name=":1">Template:Cite web</ref>

According to the type of stimulus, tropisms can be:

File:Upsidedown-tree.JPG
Example of gravitropism in the remains of a cellar of a Roman villa in the Archeologic Park in Baia, Italy

See also

References

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